Kiyik - Deer

Kiyik[1]
Vaqtinchalik diapazon: Dastlabki Oligotsen - So'nggi
Cervidae1.jpg
Cervidae oilasining bir necha a'zosining (chapdan chap tomonga soat yo'nalishi bo'yicha) tasvirlari qizil kiyik, sika kiyiklari, barasingha, kiyik, va oq dumli kiyik, va kulrang broket, va elk, va pudu.
Ilmiy tasnif e
Qirollik:Animalia
Filum:Chordata
Sinf:Sutemizuvchilar
Buyurtma:Artiodaktila
Qoidabuzarlik:Pekora
Oila:Cervidae
Goldfuss, 1820
Subfamilies
Deer range.png
Kiyiklarning barcha turlarining birlashtirilgan mahalliy turlari.

Kiyik yoki haqiqiy kiyik bor tuyoqli kavsh qaytaruvchi hayvon sutemizuvchilar shakllantirish oila Cervidae. Kiyiklarning ikkita asosiy guruhi Servinalar shu jumladan muntjac, elk (wapiti), qizil kiyik, quruq kiyik, va chital; va Capreolinae shu jumladan kiyik (karibu), kiyik, xachir kiyik, va buloq. Xitoyliklardan boshqa barcha turdagi urg'ochi kiyik va erkak kiyik suv kiyiklari, o'sib, yangi to'kib tashlang shox har yili. Bunda ular doimiy ravishda farq qiladi shoxli antilop, boshqa oilaning bir qismi bo'lgan (Bovidae ) xuddi shu tartibda juft oyoqli tuyoqlilar (Artiodactyla).

The mushk kiyik (Moschidae ) Osiyo va chevrotainlar (Tragulidae ) tropik Afrika va Osiyo o'rmonlari kavsh qaytaruvchi hayvonlar ichida alohida oilalardir (Ruminantiya ). Ular Ruminantia orasida kiyiklar bilan chambarchas bog'liq emas.

Kiyiklar san'atda paydo bo'ladi Paleolit g'or rasmlari keyin va ular bor mifologiyada rol o'ynagan, din va adabiyot tarix davomida, shuningdek geraldika, masalan, qizil kiyiklar paydo bo'ladi Olland gerbi.[2] Ularning iqtisodiy ahamiyati go'shtdan foydalanishni o'z ichiga oladi kiyik go'shti, ularning terilari yumshoq, kuchli buqa terisi va ularning shoxlari pichoq uchun tutqich sifatida. Kiyik ov qilish hech bo'lmaganda O'rta asrlardan beri ommabop faoliyat bo'lib kelgan va bugungi kunda ko'plab oilalar uchun manba bo'lib qolmoqda.

Tarqatish

Chital kiyik Nagaraxol, Hindiston

Kiyiklar turli xil yashashadi biomlar, dan tortib tundra uchun tropik tropik o'rmon. Ko'pincha o'rmonlar bilan bog'liq bo'lsa-da, ko'plab kiyiklar ekoton o'rmonlar va chakalakzorlar (qopqoq uchun) va dasht va savanna (ochiq joy) oralig'ida o'tuvchi hududlarda yashovchi turlar. Katta kiyik turlarining aksariyati dunyo bo'ylab mo''tadil aralash bargli o'rmon, tog 'aralash ignabargli o'rmon, tropik mavsumiy / quruq o'rmon va savanna yashash joylarida yashaydi. O'rmon ichidagi ochiq joylarni tozalash ma'lum darajada kiyik populyatsiyasiga ta'sir qilishi mumkin understory va kiyik eyishni yaxshi ko'radigan o'tlar, begona o'tlar va o'tlarning turlarini etishtirishga imkon berish. Bundan tashqari, qo'shni ekin maydonlariga kirish kiyiklarga ham foyda keltirishi mumkin. Biroq, aholining o'sishi va rivojlanishi uchun etarli darajada o'rmon yoki cho'tka qoplamasi ta'minlanishi kerak.

Kiyiklar keng tarqalgan bo'lib, Antarktida va Avstraliyadan tashqari barcha qit'alarda mahalliy vakillar mavjud, ammo Afrikada faqat bitta mahalliy kiyik bor, ammo Barbar stag, ning pastki turi qizil kiyik bilan cheklangan Atlas tog'lari materikning shimoli-g'arbida. Qo'shimcha yo'q bo'lib ketgan kiyik turlari, Megaceroides algericus, 6000 yil oldin Shimoliy Afrikada bo'lgan. Bug'doy Janubiy Afrikaga kiritilgan. Kichik turlari broket kiyik va pudus Markaziy va Janubiy Amerikaning, va mittjaklar Osiyo odatda zich o'rmonlarni egallaydi va ochiq joylarda kamroq uchraydi, bundan mustasno Hind muntjak. Yuqori darajada ixtisoslashgan va deyarli faqat tog'larda, o'tloqlarda, botqoqlarda va "nam" savannalarda yoki cho'llar bilan o'ralgan qirg'oq koridorlarida yashovchi kiyiklarning bir nechta turlari mavjud. Ba'zi kiyiklarning Shimoliy Amerika va Evrosiyoda sirkumpolyar tarqalishi mavjud. Bunga misollar karibu Arktik tundra va taygada (boreal o'rmonlar) va buloq yashaydigan taiga va qo'shni hududlar. Huemul kiyiklari (taruka va Chililik guemul ) Janubiy Amerikaning And ning ekologik uyalarini to'ldiring echki va yovvoyi echki, fawns echki bolalariga o'xshab o'zini tutishi bilan.

Bir suruv oq dumli kiyik, daraxtlar qoplamasi ostida harakatlanuvchi

Mo''tadil Shimoliy Amerikada yirik kiyik turlarining eng yuqori kontsentratsiyasi Kanadadagi Rokki tog'i va Kolumbiya tog'i Alberta va Britaniya Kolumbiyasi o'rtasidagi Shimoliy Amerika kiyiklarining beshta turi (oq dumli kiyik, xachir kiyik, karibu, elk, va moose) ni topish mumkin. Ushbu mintaqada bir nechta milliy bog'larning klasterlari, shu jumladan Revelstoke milliy bog'i, Glacier National Park (Kanada), Yoho milliy bog'i va Kootenay milliy bog'i Britaniya Kolumbiyasi tomonida va Banff milliy bog'i, Jasper milliy bog'i va Glacier National Park (AQSh) Alberta va Montana tomonlarida. Tog 'yonbag'iridagi yashash joylari nam ignabargli / aralashgan o'rmonli yashash joylaridan to subpine / qarag'ay o'rmonlariga qadar, alp o'tloqlari balandroq. Tog 'tizmalari orasidagi tog'oldi va daryo vodiylari ekin maydonlari va bargli park maydonlarini mozaikasi bilan ta'minlaydi. Noyob o'rmon karibu subalp yaylovlarida va undan yuqori balandlikda yashash uchun eng cheklangan hududga ega alp tundrasi ba'zi tog 'tizmalarining hududlari. Elk va xachir kiyiklari tog 'o'tloqlari va pastki ignabargli o'rmonlar orasida ko'chib yurishadi va odatda bu mintaqada keng tarqalgan. Elk, shuningdek, daryo vodiysining pastki qismida yashaydi, ular bilan oq dumli kiyiklar baham ko'rishadi. Oq quyruq so'nggi paytlarda tog'larning yon bag'irlari va daryo vodiysining pastki qismida toshlarni tog 'yonbag'irlarida o'sishga imkon beradigan bargli o'simliklarni etishtirishga imkon beradigan erlarni ekin maydonlariga aylantirish va ignabargli o'rmonlarni tozalash tufayli kengaytirdi. Ular, shuningdek, Kalgari va Edmonton shimolidagi aspen parklarida yashaydilar, u erda ular yashash joylarini mo'ylov bilan bo'lishadilar. Qo'shni Buyuk tekisliklar o'tloqlarning yashash joylari elkalar podalariga qoldirilgan, Amerika bizoni va pronghorn.

Kiyik pashshalardan saqlanish uchun qor ustida turgan podalar

The Evroosiyo Qit'a (shu jumladan Hindiston yarim oroli) dunyodagi eng ko'p kiyik turlariga ega, aksariyat turlari Osiyoda uchraydi. Evropada, taqqoslaganda, o'simlik va hayvon turlarining xilma-xilligi pastroq. Biroq, Evropadagi ko'plab milliy bog'larda va qo'riqxonalarda populyatsiya mavjud, kiyik va bug 'kiyiklari. Ushbu turlar qadimdan Evropa qit'asi bilan bog'langan, ammo ayni paytda yashaydi Kichik Osiyo, Kavkaz tog'lari va shimoli-g'arbiy Eron. Muzlik davrida "evropalik" quruq kiyiklar tarixiy jihatdan Evropaning ko'p qismida yashagan, ammo keyinchalik, asosan, hozirgi Anadolu yarim orolida cheklangan. kurka.

Hozirgi kunda Evropada quruq kiyik populyatsiyalari ushbu turni dastlab Evropaning O'rta er dengizi mintaqalariga, so'ngra Evropaning qolgan qismiga tarixiy ravishda sun'iy ravishda kiritilishining natijasidir. Dastlab ular parkdagi hayvonlar edi, keyinchalik qochib qutulishdi va yovvoyi tabiatda o'zlarini tiklashdi. Tarixiy jihatdan Evropaning kiyik turlari bargli o'rmon yashash muhitini boshqa o'txo'rlar bilan bo'lishgan, masalan, yo'q bo'lib ketgan tarpan (o'rmon oti), yo'q bo'lib ketgan Aurochs (o'rmon ho'kizlari) va yo'qolib ketish xavfi ostida aqlli (Evropa bizoni). Evropada kiyiklarni ko'rish uchun yaxshi joylarga quyidagilar kiradi Shotland tog'lari, avstriyalik Alp tog'lari, Avstriya, Vengriya va Chexiya o'rtasidagi suv-botqoq erlari va ba'zi yaxshi milliy bog'lar, shu jumladan Donana milliy bog'i Ispaniyada Veluve Gollandiyada Ardennes Belgiyada va Belowieża milliy bog'i Polsha Ispaniyada, Sharqiy Evropada va Kavkaz tog'larida hanuzgacha bokira o'rmon hududlari mavjud, ular nafaqat ko'p sonli kiyiklar populyatsiyasi, balki ilgari donishmandlar kabi ko'p bo'lgan boshqa hayvonlar ham yashaydi. Evroosiyo lyuksi, Iberian lynx, bo'rilar va jigarrang ayiqlar.

Mo''tadil Osiyoda yirik kiyik turlarining eng yuqori kontsentratsiyasi aralash bargli o'rmonlarda, tog 'ignabargli o'rmonlarida va Taygada Shimoliy Koreya, Manchuriya (Shimoliy-Sharqiy Xitoy) va Ussuri viloyati (Rossiya) bilan chegaradosh joylarda uchraydi. Bular dunyodagi eng boy bargli va ignabargli o'rmonlardan biridir Sibir kiyiklari, sika kiyiklari, elk va buqalar. Osiyo karibulari ushbu mintaqaning shimoliy chekkalarini Xitoy-Rossiya chegarasi bo'ylab egallaydi.

Sika kiyiklari kabi kiyiklar, Toroldning kiyiklari, O'rta Osiyo qizil kiyiklari va elklar tarixiy ravishda shoxlari uchun parvarish qilingan Xan xitoylari, Turkiy xalqlar, Tungus xalqlari, Mo'g'ullar va Koreyslar. Kabi Sami xalqi Finlyandiya va Skandinaviya, tungus xalqlari, mo'g'ullar va Janubiy Sibir, Shimoliy Mo'g'uliston va Ussuri mintaqasidagi turkiy xalqlar ham Osiyo karibularining yarim uyli podalarini boqishga kirishdilar.

Tropikada yirik kiyik turlarining eng yuqori kontsentratsiyasi Janubiy Osiyoda Hindistonning Hind-Gangetik tekisligi va Nepalning Teray mintaqalarida uchraydi. Ushbu serhosil tekisliklar tropik mavsumiy namli bargli, quruq bargli o'rmonlardan va quruq va ho'l savannalardan iborat. chital, cho'chqa kiyiklari, barasingha, Hind sambar va Hind muntjak. Yo'qolib ketish xavfi ostida bo'lgan barasingha va juda keng tarqalgan chital kabi o'tlatish turlari ochko'z bo'lib, katta podalarda yashaydi. Hind sambarasi ochko'z bo'lishi mumkin, lekin odatda yolg'iz yoki kichikroq podalarda yashaydi. Cho'chqa kiyiklari yakka va zichligi hindu mintjakiga qaraganda pastroq. Kiyiklarni Hindiston, Nepal va Shri-Lankadagi bir nechta milliy bog'larda ko'rish mumkin Kanha milliy bog'i, Dudhva milliy bog'i va Chitvan milliy bog'i eng mashhur. Shri-Lankaning Wilpattu milliy bog'i va Yala milliy bog'i hind sambar va chitalning katta podalariga ega. Hindiston sambarasi Shri-Lankada boshqa hududlarga qaraganda ko'proq ochko'zroq va boshqa joylarga qaraganda ko'proq podalarni shakllantirishga moyil.

Tailandning Chao Praya daryosi vodiysi bir paytlar asosan yo'q bo'lib ketgan cho'chqa kiyiklari populyatsiyasini joylashtirgan tropik mavsumiy nam bargli o'rmon va ho'l savanna bo'lgan. Schomburgk kiyiklari, Eldning kiyiklari, Hind sambarasi va hindu muntjak. Cho'chqa kiyiklari ham, Eldning kiyiklari ham kamdan-kam uchraydi, hind sambarlari va hindu muntjaklari qo'riqlanadigan milliy bog'larda, masalan, gullab-yashnaydilar. Khao Yai. Ushbu Janubiy Osiyo va Janubi-Sharqiy Osiyo kiyik turlarining aksariyati yashash muhitini boshqalar bilan bo'lishadi o'txo'rlar, kabi Osiyo fillari, turli xil Osiyo karkidon turlari, turli antilopalar turlari (masalan.) nilgay, to'rt shoxli antilop, Blackbuck va Hind jayroni Hindistonda) va yovvoyi buqalar (masalan yovvoyi Osiyo suvi buvali, gaur, banteng va kuprey ). Muayyan hududda turli xil o'txo'rlar birgalikda yashashi mumkin bo'lgan usullardan biri shundaki, har bir tur turli xil oziq-ovqat imtiyozlariga ega, garchi bir-birining ustiga chiqadigan bo'lsa ham.

Avstraliyada oltitasi bor kiritilgan turlar barqaror yovvoyi populyatsiyasini tashkil etgan kiyiklarning akklimatizatsiya jamiyati 19-asrda chiqarilgan nashrlar. Bular quruq bug ', qizil kiyik, sambar, cho'chqa kiyiklari, rusa va chital. 1851 yilda ingliz va shotland zaxiralaridan Yangi Zelandiyaga olib kirilgan qizil kiyiklar uy sharoitiga keltirildi kiyik fermer xo'jaliklari 1960 yillarning oxiriga kelib, hozirgi kunda u erda oddiy qishloq xo'jalik hayvonlari mavjud. Yangi Zelandiyada yana etti turdagi kiyiklar paydo bo'ldi, ammo ularning hech biri qizil kiyik kabi keng tarqalmagan.[3]

Tavsif

Kiyiklar artiodaktilalarning bovidlardan keyin ikkinchi turkumini tashkil qiladi.[4] Shunga o'xshash tuzilishga ega bo'lsa ham, kiyiklar juda ajralib turadi antilopalar ular tomonidan shox, doimiydan farqli o'laroq vaqtincha va muntazam ravishda o'sib boradi shoxlar qovurg'a.[5] Kiyiklarga xos xususiyatlarga uzun, kuchli oyoqlar, kichraytiruvchi quyruq va uzun quloqlar kiradi.[6] Kiyiklar jismoniy nisbatlarda keng farq qiladi. Hozirgacha mavjud bo'lgan eng katta kiyik buloq bo'yi qariyb 2,6 metr (8,5 fut) va vazni 800 kilogrammgacha (1800 funt).[7][8] Elk elkasida 1,4–2 metr (4,6-6,6 fut) turadi va vazni 240–450 kilogramm (530–990 funt).[9] Shimoliy pudu - bu dunyodagi eng kichik kiyik; u yelkasida 32-35 santimetrga (13-14 dyuym) etadi va 3.3-6 kilogrammni (7.3-13.2 funt) tashkil etadi. Janubiy pudu biroz balandroq va og'irroq.[10] Jinsiy dimorfizm juda aniq - aksariyat turlarda erkaklar urg'ochilarnikidan kattaroq,[11] va kiyikdan tashqari, faqat erkaklar shoxga ega.[12]

Palto rangi odatda qizil va jigarrang o'rtasida farq qiladi,[13] ammo shoxli kiyikda shokolad jigarrang kabi qorong'i bo'lishi mumkin[14] yoki elkada bo'lgani kabi kulrang rangga ega.[9] Broket kiyiklarining turli xil turlari palto rangidan kul rangdan qizil jigar ranggacha farq qiladi.[15] Chital kabi bir necha tur,[16] tushgan kiyik[17] va sika kiyiklari[18] jigarrang paltosidagi oq dog'lar. Kiyik kiyimi sezilarli geografik o'zgarishni namoyish etadi.[19] Kiyiklar ikkitadan o'tishadi naychalar bir yilda;[13][20] Masalan, qizil kiyikda qizil, ingichka sochli yozgi palto asta-sekin kuzda zich, kulrang jigarrang qishki paltos bilan almashtiriladi, bu esa keyingi bahorda yozgi paltosga yo'l beradi.[21] Moulting ta'sir qiladi fotoperiod.[22]

Kiyiklar ham zo'r saklovchilar va suzuvchilar. Kiyiklar kavsh qaytaruvchi hayvonlar, yoki to'rtta kamerali oshqozonga ega. Ba'zi bir kiyiklar, masalan, orolda bo'lganlar ROM,[23] mavjud bo'lganda go'sht iste'mol qiling.[24]

Yalang'ochning birinchi qadamlari

Deyarli barcha kiyiklarning har bir ko'z oldida yuz bezi bor. Bezda kuchli hid mavjud feromon, odatlangan belgi uning uy oralig'i. Turli xil turdagi buklar bu bezlarni g'azablanganda yoki hayajonlanganda keng ochadi. Barcha kiyiklarda a jigar a .siz o't pufagi. Kiyiklarda ham bor tapetum lucidum, bu ularga etarlicha yaxshi narsalarni beradi tungi ko'rish.

Shoxlar

Ikki Sambar kiyik jang qilish, Silvassa, Hindiston

Hamma erkaklar kiyiklari bor shox, bundan mustasno suv kiyiklari, unda erkaklar pastki jag 'ostiga etib boradigan uzun tusga o'xshash itlarga ega.[25] Odatda urg'ochilarda shox yo'q, ammo ayol kiyik kiyiklari erkaklarnikiga qaraganda kichikroq va kam tarvaqaylab qo'ygan.[26] Ba'zan boshqa turlardagi urg'ochi shoxlar paydo bo'lishi mumkin, ayniqsa, Evropa kiyiklari, qizil kiyiklar, oq dumaloq va xachir kiyiklari va kamroq plesiometakarpal kiyiklari kabi telemetakarpal kiyiklarida. Shoxli oq dumli kiyikni o'rganish shuni ko'rsatdiki, shoxlar mayda va yomon shaklga ega bo'lib, tug'ruq paytida tez-tez to'kiladi.[27]

Bug'u kiyiklari va kiyikning turli xil kichik turlari eng katta va eng og'ir shoxlarga ega, ham mutloq ma'noda, ham tana massasiga mutanosib ravishda (tana vaznining har bir kilogrammiga o'rtacha 8 gramm (0,28 oz));[26][28] tukli kiyiklarda esa barcha kiyiklarning eng kichkina shoxlari bor, pududa esa tana massasiga nisbatan eng engil shoxlar mavjud (tana vazniga kilogramm uchun 0,6 gramm (0,021 oz)).[26] Shoxlarning tuzilishi sezilarli darajada o'zgarib turadi; kiyik va elk shoxlari palma (markaziy qismi keng) bo'lsa, oq quyruq kiyiklari shoxlari oldinga egilgan asosiy nurdan yuqoriga chiqadigan bir qator tishlarni o'z ichiga oladi, pudu esa shunchaki boshoqdir.[10] Shoxning rivojlanishi hayvonning bir yoshga to'lguniga qadar bosh suyagining tepasida paydo bo'ladigan suyak tuzilishi bo'lgan pedikeldan boshlanadi. Pedicel keyingi yil tikanli shoxni keltirib chiqaradi, uning o'rniga uchinchi yilda tarvaqaylab qo'yilgan shox bilan almashtiriladi. Kattaroq va tarvaqaylab to'plamni yaratish uchun shoxlar to'plamini yo'qotish jarayoni butun hayot davomida davom etadi.[26] Shoxlar yumshoq bo'lib chiqadi to'qimalar (baxmal shoxlari deb nomlanuvchi) va asta-sekin suyak tuzilmalariga (qattiq shoxlar deb nomlanuvchi) aylanib qattiqlashadi mineralizatsiya va blokirovka qon tomirlari to'qimada, uchidan poydevorigacha.[29]

Antlers eng abartılı erkaklardan biri bo'lishi mumkin ikkilamchi jinsiy xususiyatlar,[30] va asosan reproduktiv muvaffaqiyatga erishish uchun mo'ljallangan jinsiy tanlov va jang uchun. Shox ustidagi tishlar (vilkalar) boshqa erkaklar shoxlarini joyiga qulflashga imkon beradigan oluklar hosil qiladi. Bu erkaklar yuziga shikast etkazmasdan kurashishga imkon beradi.[31] Antlers shaxsning ijtimoiy ierarxiyadagi pozitsiyasi va uning xulq-atvori bilan o'zaro bog'liqdir. Masalan, shoxlar qanchalik og'ir bo'lsa, shaxsning ijtimoiy ierarxiyadagi mavqei shunchalik yuqori bo'ladi va shoxlarning to'kilishi shunchalik kechikadi;[26] katta shoxli erkaklar boshqalarga nisbatan ko'proq tajovuzkor va hukmron bo'lish xususiyatiga ega.[32] Antlers bo'lishi mumkin halol signal genetik sifat; tana kattaligiga nisbatan kattaroq shoxli erkaklar qarshilikni kuchaytiradi patogenlar[33] va reproduktiv salohiyat yuqori.[34]

Elkda Yellowstone milliy bog'i, shoxlar, shuningdek, yirtqich hayvonlardan himoya qiladi bo'rilar.[35]

Tishlarning gomologiyasi, ya'ni turlar orasida shoxlarning tarvaqaylab tuzilishi 1900-yillarga qadar muhokama qilingan.[36][37][38] Yaqinda shoxlarning tarvaqaylab tuzilishini tavsiflash va tishlarning homologiyasini aniqlashning yangi usuli ishlab chiqildi.[39]

Tishlar

Kiyiklarning aksariyati 32 tishga ega; tegishli tish formulasi bu: 0.0.3.33.1.3.3. Elk va kiyik istisno holatlar bo'lishi mumkin, chunki ular yuqori tishlarini ushlab turishi va shu bilan 34 tishiga ega bo'lishi mumkin (tish formulasi: 0.1.3.33.1.3.3).[40] Xitoy suv kiyiklari, tutuk kiyiklari va muntjac yuqori qismi kattalashtirilgan it tishlari o'tkir tishlarni hosil qiladi, boshqa turlarda esa ko'pincha yuqori itlar etishmaydi. Kiyikning yonoq tishlari emalning yarim oylik tizmalariga ega bo'lib, ular turli xil o'simliklarni maydalashga imkon beradi.[41] Kiyiklarning tishlari o'simlik bilan oziqlanishga moslashgan va boshqa kavsh qaytaruvchi hayvonlar singari ularning yuqori qismi yo'q tish kesuvchi, buning o'rniga yuqori jag'ning old qismida qattiq yostiq bor.

Biologiya

Fawn

Parhez

Kiyiklar brauzerlar va asosan barglar bilan oziqlaning o'tlar, toshlar, forblar, butalar va daraxtlar, qo'shimcha iste'mol bilan likenler qish paytida shimoliy kengliklarda.[42] Ular tomonidan kichik, ixtisoslashtirilmagan oshqozon bor kavsh qaytaruvchi hayvon standartlar va yuqori ovqatlanish talablari. Masalan, ko'p miqdordagi tolali ovqatni iste'mol qilish va hazm qilish o'rniga, masalan, qo'ylar va qoramol qil, kiyik oson hazm bo'ladigan kurtaklar, yosh barglar, yangi o'tlar, yumshoq novdalar, mevalar, qo'ziqorinlar va likenler. Kam tolali oziq-ovqat, minimal fermentatsiya va parchalanishdan so'ng, oziq-ovqat kanalidan tez o'tadi. Kiyik kabi ko'plab minerallarni talab qiladi kaltsiy shox o'sishini qo'llab-quvvatlash maqsadida fosfat va bu qo'shimcha ravishda ozuqaviy moddalarga boy parhezni talab qiladi. Shu bilan birga, kiyiklarning go'shtli faoliyat bilan shug'ullanishi, masalan, o'lik yeyish kabi ba'zi xabarlari mavjud xotinlar ko'l bo'ylarida[43] yoki uyalarini kamsitish shimoliy bobvitlar.[44]

Ko'paytirish

Ayol elk emizuvchi yosh

Deyarli barcha servikslar deyiladi uniparental turlar: qoraqalpog'istonni faqat kapalak deb nomlanuvchi ona parvarish qiladi. Odatda, kapalakda bir vaqtning o'zida bitta yoki ikkita sichqonchani bo'ladi (uch egizaklar, noma'lum bo'lsa ham, kamdan-kam uchraydi). Juftlik mavsumi odatda avgust oyining oxirida boshlanadi va dekabrgacha davom etadi. Ba'zi turlar mart oyining boshiga qadar juftlashadi. The homiladorlik davri Evropada kiyik uchun o'n oygacha bo'lgan joyda. Fawnlarning aksariyati mo'ynalari oq dog'lar bilan o'ralgan holda tug'iladi, garchi ko'plab turlarda ular birinchi qishining oxiriga kelib bu joylarni yo'qotadilar. Yalang'och hayotning dastlabki yigirma daqiqasida yirtqich birinchi qadamlarini qo'yishni boshlaydi. Onasi uni deyarli hiddan xoli bo'lmaguncha tozalaydi, shuning uchun yirtqichlar uni topa olmaydi. Onasi tez-tez boqish uchun ketib qoladi, jo'ja esa ortda qolishni yoqtirmaydi. Ba'zida onasi uni oyog'i bilan muloyimlik bilan pastga itarishi kerak.[45] Yalang'och onasi bilan yurishga qodir bo'lguncha bir hafta davomida maysada yashirin qoladi. Yalang'och va uning onasi taxminan bir yil birga bo'lishadi. Erkak odatda tashlab ketadi va onasini boshqa ko'rmaydi, lekin urg'ochilar ba'zan o'zlarining sansarlari bilan qaytib kelib, kichik podalarni hosil qiladilar.

Kasallik

Buyuk Britaniyaning ba'zi hududlarida kiyiklar (ayniqsa quruq kiyik ularning tufayli g'ayrioddiy xatti-harakatlar ), uzatilishi mumkin bo'lgan suv ombori sifatida ko'rsatilgan sigirning sil kasalligi,[46][47] bu kasallik 2005 yilda Buyuk Britaniyada uni yo'q qilishga urinish uchun 90 million funt sterlingga tushgan.[48] Yangi Zelandiyada kiyiklar vektorlarni yig'ishda muhim ahamiyatga ega deb o'ylashadi M. bovis brushtail egalik qiladigan joylarda Trichosurus vulpecula yuqtirgan va tana go'shti boshqa joylarda tozalanganida, uni ilgari yuqtirilmagan xayvonlarga o'tkazgan.[49] Oq quyruq Odocoileus virginianus Michigan shtatidagi sigir tüberkülozunun yagona xizmat ko'rsatuvchisi sifatida tasdiqlangan, bu AQShda chorvachilikda kasallikni butun mamlakat bo'ylab yo'q qilish uchun muhim to'siq bo'lib qolmoqda.[50]

Bug'u va kiyik ko'tarishi mumkin quturish.[51]

Yurak mushuklari azoblanishi mumkin miya qurti, a gelmint u tuxum qo'yishi uchun mos joy qidirishda miya orqali teshiklarni ochadi. Hukumat biologining ta'kidlashicha, "Ular kerakli joyni qidirib yurishadi va hech qachon uni topa olmaydilar". Kiyiklar bu parazitga qarshi immunitetga ega ko'rinadi; u ovqat hazm qilish tizimidan o'tadi va najas bilan ajralib chiqadi. Parazit mo'ylov ichaklari tomonidan skrining qilinmaydi va miyaga o'tadi, u erda xatti-harakatlarida ham, yurishida ham tashqi ko'rinishidan zarar ko'riladi.[51]

Kiyik, elk va buqalar Shimoliy Amerika azoblanishi mumkin surunkali isrof kasalligi, da aniqlangan Kolorado 1960-yillarda laboratoriya va prion kasalligi deb ishoniladi. Ehtiyotkorlik tufayli ovchilarga aloqa qilishdan qochish tavsiya etiladi ko'rsatilgan xavf materiallari (SRM), masalan, miya, o'murtqa ustun yoki limfa tugunlari. Pichoq va so'yish uchun ishlatiladigan boshqa asboblarni so'yishda va dezinfektsiyalashda go'shtni debonatsiyalash hukumatning boshqa tavsiyalaridan biridir.[52]

Evolyutsiya

Kiyiklar shoxsiz rivojlangan, tusked zamonaviyga o'xshash ajdodlar duikerlar va kichkina kiyiklar boshida Eosen va asta-sekin rivojlanib, birinchi antlered servoidlarga ( superfamily tservidlar va unga aloqador qirilib ketgan oilalar) da Miosen. Oxir-oqibat, shoxlarning rivojlanishi bilan, shuningdek, yuqori qismlar tish kesuvchi ko'zdan yo'qoldi. Shunday qilib, kiyiklarning rivojlanishi taxminan 30 million yil davom etdi. Biolog Valerius Geyst evolyutsiyani bosqichma-bosqich sodir bo'lganligini ko'rsatadi. Ushbu evolyutsiyani kuzatish uchun juda ko'p taniqli qazilma qoldiqlari mavjud emas, ammo faqat skervit va shox parchalari bo'lib, ular osongina servid bo'lmagan turlarning soxta shoxlari bilan aralashishi mumkin.[10][53]

Eosen davri

The kavsh qaytaruvchi hayvonlar, Cervidae ajdodlari, kelib chiqqan deb ishoniladi Diakodeksis, ma'lum bo'lgan eng qadimgi artiodaktil (juft oyoqli tuyoqlilar), Eosendagi 50-55 Mya.[54] Diakodeksis, deyarli a kattaligi quyon, xususiyatli suyak suyagi barcha zamonaviylarga xos juft oyoqli tuyoqlilar. Ushbu ajdod va uning qarindoshlari butun Shimoliy Amerika va Evroosiyoda sodir bo'lgan, ammo kamida 46 Mya tomonidan pasayib ketgan.[54][55] Ning deyarli to'liq skeletini tahlil qilish Diakodeksis 1982 yilda kashf etilgan bu ajdod kavsh qaytaradigan hayvonlarga qaraganda kavsh qaytaruvchi hayvonlarga yaqinroq bo'lishi mumkin degan taxminlarni keltirib chiqardi.[56] Andromeriks prehistorik yana bir taniqli kavsh qaytaruvchi hayvondir, ammo unga yaqinroq ko'rinadi tragulidlar.[57]

Oligotsen davri

Ning shakllanishi Himoloy va Alp tog'lari muhim geografik o'zgarishlarni keltirib chiqardi. Bu kiyikka o'xshash shakllarning keng xilma-xilligi va servitlarning paydo bo'lishining asosiy sababi edi Oligotsen erta Plyotsen.[58] Oligotsenning ikkinchi yarmi (28-34 Mya) Evropaning ko'rinishini ko'rdi Eumeryx va Shimoliy Amerika Leptomeriks. Ikkinchisi tish morfologiyasida zamonaviy bovid va servidlarga o'xshardi (masalan, shunday edi) brachyodont oldingi tish ko'proq bo'lgan bo'lsa) rivojlangan.[59] Kiyikka o'xshash boshqa shakllarga Shimoliy Amerika ham kiradi Blastomeriks va Evropa Dremoteliy; bu shilimshiq tishli hayvonlar barcha zamonaviy shoxli kiyiklarning bevosita ajdodlari bo'lgan deb hisoblashadi, garchi o'zlarida shox yo'q edi.[60] Yana bir zamonaviy shakl to'rt shoxli edi protokeratid Protokeralar, bu bilan almashtirildi Sindiyokeralar miosenda; bu hayvonlar burunda shox borligi bilan noyob bo'lgan.[53] Shimoliy Amerikada topilgan, taxminan 35 million yil avvalgi so'nggi Eosen qoldiqlari buni ko'rsatadi Sindiyokeralar Boshsuyagi suyaksiz bo'lib, bargsiz shoxga o'xshash edi.[61]

Miosen davri

Qadimgi topilmalar shuni ko'rsatadiki, Cervoidea superfamilasining eng qadimgi a'zolari Eviosiyoda Miosenda paydo bo'lgan. Dikrocerus, Evropoks va Heteroproks ehtimol birinchi shoxli servikslar bo'lgan.[62] Dikrocerus muntazam ravishda to'kib tashlanadigan bitta vilka shoxlarini namoyish etdi.[63] Stefanokemalar ko'proq rivojlangan va tarqoq ("tojli") shoxlarga ega edi.[64] Procervulus (Palaeomerycidae ) ning tishlariga qo'shimcha ravishda Dremoteliy, egiluvchan shoxlar to'kilmagan.[65] Kabi zamonaviy shakllar merikodontinlar pirovardida zamonaviy pronghorn paydo bo'ldi.[66]

Servinalar 7-9 Mya atrofida, Markaziy Osiyoda kech bo'lgan Miosen davrida, mavjud bo'lgan servikslarning birinchi guruhi sifatida paydo bo'ldi. Muntiacini qabilasi tashqi ko'rinishini shunday qildi Muntiacus leilaoensis 7-8 Mya atrofida;[67] Dastlabki maxtaklarning o'lchamlari har xil edi - quyonlar kabi kichkina yoki bug 'kiyiklari kabi katta. Ularning jang qilish uchun tishlari va himoya qilish uchun shoxlari bor edi.[10] Tez orada Capreolinae ergashdi; Alceini 6.4-8.4 Mya paydo bo'ldi.[68] Taxminan shu davrda Tetis okeani keng yaylovlarga yo'l berish uchun g'oyib bo'ldi; bular kiyiklarga mo'l-ko'l oqsillarga boy o'simliklarni taqdim etdi, bu esa bezak shoxlarini rivojlanishiga olib keldi va populyatsiyalarning gullab-yashnashi va hududlarni mustamlaka qilishiga imkon berdi.[10][58] Shoxlar talaffuz qilinishi bilanoq, itlar saqlanib qolmagan yoki yomon namoyish qilingan (masalan, elkada bo'lgani kabi), ehtimol diet endi yo'q edi ko'rib chiqing - hukmronlik qilgan va shoxli organlar yaxshiroq edi. Muntjak va tukli kiyiklarda shoxlar bilan bir qatorda itlar ham kichkina. Ammo tragulidlar bugungi kungacha uzun tishlarga ega.[55]

Pliyotsen davri

Ning boshlanishi bilan Plyotsen, global iqlim salqinlashdi. Dengiz sathining qulashi katta muzliklarga olib keldi; binobarin, o'tloqlar to'yimli em-xashaklarga boy bo'ldi. Shunday qilib, kiyik populyatsiyasida yangi shovqin paydo bo'ldi.[10][58] Cervini-ning eng keksa a'zosi, Servocerus novorossiae, Evtosiyoning miosendan pliosenga (4.2-6 Mya) o'tish davrida paydo bo'lgan;[69] servitsin qoldiqlari Plyotsenning boshidan to kechgacha Pleystotsen Xitoyda qazilgan[70] va Himoloy.[71] Esa Servus va Dama deyarli 3 Mya paydo bo'ldi, Eksa pliyotsen-pleistosen davrida paydo bo'lgan. Kapreolini va Rangiferini qabilalari 4-7 Mya atrofida paydo bo'lgan.[68]

Taxminan 5 Mya, rang oralig'i Bretziya va Eocoileus Shimoliy Amerikaga etib kelgan birinchi servidlar edi.[68] Bu shuni anglatadiki, Bering bo'g'ozini kechgi miosen-plyotsen davrida kesib o'tish mumkin; bu juda katta ehtimollik bilan ko'rinadi tuyalar bir vaqtning o'zida Shimoliy Amerikadan Osiyoga ko'chib kelgan.[72] Kiyik Janubiy Amerikani Pliyosenning oxirida (2,5–3 Mya) bosib olgan Buyuk Amerika almashinuvi, yaqinda tashkil etilganiga rahmat Panama Istmusi, va qit'ada raqobatlashadigan kavsh qaytaruvchi hayvonlar kamligi tufayli muvaffaqiyatli paydo bo'ldi.[73]

Pleystotsen davri

Ta'sirchan shoxlari bo'lgan katta kiyiklar, pleystotsen davrida, ehtimol evolyutsiyani qo'zg'atish uchun mo'l-ko'l resurslar natijasida rivojlangan.[10] Dastlabki pleystotsen servervidi Evkladokeros hajmi bo'yicha zamonaviy elk bilan solishtirish mumkin edi.[74] Megaloceros (Pliyotsen-Pleystosen) tarkibida Irlandiyalik elk (M. giganteus), ma'lum bo'lgan eng katta servikslardan biri. Irlandiyalik elk yelkasida 2 metrga (6,6 fut) yetdi va og'ir shoxlari bor edi, ular uchidan uchigacha 3,6 metrni tashkil qildilar.[75] Ushbu yirik hayvonlar o'rtasidagi ziddiyat tufayli yo'q bo'lib ketishga duch kelgan deb o'ylashadi jinsiy tanlov katta shox va tana uchun va tabiiy selektsiya kichikroq shakl uchun.[76] Ayni paytda, ariq va kiyik Shimoliy Amerikaga Sibirdan tarqaldi.[77]

Taksonomiya va tasnif

Servidning bosh suyagi

Kiyiklar artiodaktil oila Cervidae. Bu oila birinchi edi tasvirlangan nemis zoologi tomonidan Georg August Goldfuss yilda Handbuch der Zoologie (1820). Uch subfamilies tanilgan: Capreolinae (birinchi bo'lib ingliz zoologi tomonidan tasvirlangan Joshua Bruks Cervinae (Goldfuss tomonidan tasvirlangan) va Hydropotinae (birinchi bo'lib frantsuz zoologi tomonidan tasvirlangan) Eduard Lui Troyessart 1898 yilda).[4][78]

Kiyiklarni tasniflash bo'yicha boshqa urinishlar morfologik va genetik farqlar.[53] Angliya-irland tabiatshunosi Viktor Bruk 1878 yilda kiyiklarni ikkinchi va beshinchi xususiyatlariga ko'ra ikki sinfga bo'lish mumkin degan taklif metakarpal suyaklar ularning oldingi oyoqlari: Plesiometakarpaliya (eng qadimgi dunyo kiyiklari) va Telemetakarpaliya (eng yangi dunyo kiyiklari). U davolagan mushk kiyik Servid sifatida, uni Telemetakarpaliya ostiga qo'yadi. Telemetakarpal kiyik faqat bo'g'imdan uzoqda joylashgan elementlarni ko'rsatgan bo'lsa, plesiometakarpal kiyik qo'shimchaga yaqin elementlarni ham saqlab qoldi.[79] Asosida farqlash diploid soni xromosomalar 20-asrning oxirlarida bir nechta nomuvofiqliklar tufayli kamchiliklarga yo'l qo'yildi.[53]

1987 yilda zoologlar Kolin Groves va Piter Grubb uchta subfamilani aniqladi: Cervinae, Hydropotinae va Odocoileinae; ular gidropotinlarda shox yo'qligini, qolgan ikki subfamiliya esa skelet morfologiyasi bilan ajralib turishini ta'kidladilar.[80] Biroq, ular 2000 yilda ushbu tasnifdan qaytishdi.[81]

Tashqi aloqalar

21-asrning boshiga qadar bu oila deb tushunilgan Moschidae (mushk kiyik) opa Cervidae-ga. Biroq, 2003 yil filogenetik Aleksandr Xasaninning tadqiqotlari (ning Milliy tabiiy tarix muzeyi, Frantsiya ) va hamkasblar, asoslangan mitoxondrial va yadroviy tahlillari, Moschidae va Bovidae shakl qoplama Cervidae singlisi. Tadqiqotga ko'ra, Cervidae ajratilgan 27-28 million yil oldin Bovidae-Moschidae to'shagidan.[82] Quyidagi kladogramma 2003 yilgi tadqiqotga asoslangan.[82]

Ruminantiya
Tragulina

Tragulidae Kantschil-drawing white background.jpg

Pekora

Antilokapridae Antilocapra white background.jpg

Giraffidae Giraffa camelopardalis Brockhaus white background.jpg

Cervidae The deer of all lands (1898) Hangul white background.png

Bovidae Birds and nature (1901) (14562088237) white background.jpg

Moschidae Moschus chrysogaster white background.jpg

Ichki munosabatlar

2006 yil filogenetik Cervidae-dagi ichki munosabatlarni Klement Gilbert va uning hamkasblari tomonidan o'rganish oilani ikkita yirik to'qnashuvga ajratdi: Capreolinae (telemetakarpal yoki yangi dunyo kiyiklari) va servinalar (plesiometakarpal yoki eski dunyo kiyiklari). 20-asrning oxiridagi tadqiqotlar oilada shunga o'xshash bifurkatsiyani taklif qildi. Bu va avvalgi tadqiqotlar qo'llab-quvvatlanmoqda monofil Cervinae-da, Capreolinae paydo bo'lganda parafiletik. 2006 yilgi tadqiqot Cervinae-da, Cervini-da (avlodni o'z ichiga olgan) ikkita naslni aniqladi Eksa, Servus, Dama va Rucervus ) va Muntiacini (Muntiakus va Elafod ). Capreolinae uchta naslni namoyish etdi, Alceini (Alces turlari), Kapreolini (Kapreol va Hydropotinae subfamily) va Rangiferini (Blastokerus, Gipokamel, Mazama, Odocoileus, Pudu va Rangifer turlari). Quyidagi kladogramma 2006 yilgi tadqiqotga asoslangan.[68]

Cervidae
Servinalar
Muntiacini

Rivzning munajjagi

Tugilgan kiyik The deer of all lands (1898) Michie's tufted deer white background.png

Cervini

Bug'doy Atlas de poche des mammifères de France, de la Suisse romane et de la Belgique (Pl. 43) (Dama dama).jpg

Fors kiyiklari

RusaThe deer of all lands (1898) Moluccan rusa white background.png

Sambar The deer of all lands (1898) Indian sambar white background.png

Qizil kiyik The deer of all lands (1898) Hangul white background.png

Toroldning kiyiklariThe deer of all lands (1898) Thorold's deer white background.png

Sika kiyiklariThe deer of all lands (1898) Manchurian sika white background.png

Elk

Eldning kiyiklariThe deer of all lands (1898) Siamese thameng white background.png

Pere Devidning kiyiklariElaphurusdavidianus white background.jpg

BarasinghaThe deer of all lands (1898) Swamp deer white background.png

Hind cho'chqa kiyiklari

ChitalThe deer of all lands (1898) Chital white background.png

Capreolinae
Rangiferini

Kiyik (Karibu) The deer of all lands (1898) Scandinavian reindeer white background.png

Amerika qizil broketi PZSL1850PlateMammalia24 Mazama americana.png

Oq dumli kiyik The deer of all lands (1898) Virginia deer white background.png

Xachir kiyik The deer of all lands (1898) Mule deer white background.png

Marsh kiyik

Kulrang braket

Janubiy pudu Pudu puda Werner (white background).JPG

Taruka The deer of all lands (1898) Peruvian guemal white background.png

Kapreolini

Kiyik The deer of all lands (1898) European roe deer white background.png

Suv kiyiklari The deer of all lands (1898) Chinese water deer white background.png

Alceini

Mus yoki Evroosiyo elkasi The deer of all lands (1898) Elk white background.png

Mavjud subfamilalar, avlodlar va turlar

Subfamila Capreolinae 9 avlod va 36 turdan iborat, shu bilan birga Servinalar 10 turkum va 55 turdan iborat.[78][tekshirib bo'lmadi ] Gidropotinalar bitta turdan iborat, ya'ni suv kiyiklari (H. inermis); ammo, 1998 yildagi bir tadqiqot uni Capreolinae-ning ostiga qo'ydi.[83] Quyidagi ro'yxat Groves va Grubb kabi zoologlarning molekulyar va filogenetik tadqiqotlariga asoslangan.[84][85][86][87][88]

Extinct subfamilies, genera and species

The following is the classification of the extinct cervids with known fossil record:

Insonlarning o'zaro ta'siri

Yuqori paleolit cave painting a Megaloceros giant deer at Lascaux, 17,300 years old

In prehistory

Deer were an important source of food for early hominids. Xitoyda, Homo erectus fed upon the sika deer, while the red deer was hunted in Germany. In Yuqori paleolit, the reindeer was the staple food for Kromagnon odamlar,[107] esa g'or rasmlari da Lascaux in southwestern France include some 90 images of stags.[108]

Tarixda

Qadimgi yunoncha gilt-silver riton, 4th century BC

Deer had a central role in the ancient art, culture and mythology of the Xettlar, qadimgi misrliklar, Keltlar, qadimgi yunonlar, the Asians and several others. Masalan, Stag Hunt Mosaic qadimiy Pella, ostida Makedoniya qirolligi (4th century BC), possibly depicts Buyuk Aleksandr hunting a deer with Hephaistion.[109] Yapon tilida Shintoism, the sika deer is believed to be a messenger to the gods. In China, deer are associated with great medicinal significance; deer penis is thought by some in China to have afrodizyak xususiyatlari.[110] Spotted deer are believed in China to accompany the god of longevity. Deer was the principal sacrificial animal for the Huichal Indians of Mexico. In medieval Europe, deer appeared in hunting scenes and coats-of-arms. Deer are depicted in many materials by various pre-Hispanic civilizations in the Andes.[107][111]

The common male first name Oskar is taken from the Irland tili, where it is derived from two elements: the first, os, means "deer"; the second element, cara, means "friend". The name is borne by a famous hero of Irlandiya mifologiyasiOskar, nabirasi Fionn Mac Cumhail. The name was popularised in the 18th century by James Macpherson, creator of 'Ossianic poetry'.

Adabiyotda

In the Indian epic Ramayana, Sita is lured by a golden deer (maricha)

Deer have been an integral part of fables and other literary works since the inception of writing. Stags were used as symbols in the latter Sumerian writings. For instance, the boat of Sumerian god Enki is named the Stag of Azbu. There are several mentions of the animal in the Rigveda shuningdek Injil. In the Indian epic Ramayana, Sita is lured by a golden deer which Rama tries to catch. In the absence of both Rama and Lakshman, Ravana kidnaps Sita. Many of the allegorical Aesop's fables, such as "The Stag at the Pool", "The One-Eyed Doe" and "The Stag and a Lion", personify deer to give moral lessons. For instance, "The Sick Stag" gives the message that uncaring friends can do more harm than good.[107] The Yaqui deer song accompanies the deer dance which is performed by a pascola [from the Spanish 'pascua', Easter] dancer (also known as a deer dancer). Pascolas would perform at religious and social functions many times of the year, especially during Lent and Easter.[107][112]

In one of Rudolf Erich Raspe 's 1785 stories of Baron Münxauzen 's Narrative of his Marvellous Travels and Campaigns in Russia, the baron encounters a stag while eating cherries and, without ammunition, fires the cherry-pits at the stag with his musket, but it escapes. The next year, the baron encounters a stag with a cherry tree growing from its head; presumably this is the animal he had shot at the previous year. Yilda Rojdestvo lore (such as in the narrative poem "Aziz Nikolayga tashrif "), kiyik are often depicted pulling the chana ning qor bobo, Santa Klaus.[113] Marjorie Kinnan Rawlings "s Pulitser mukofoti -winning 1938 novel Yilboshi was about a boy's relationship with a baby deer. The fiction book Fire Bringer is about a young fawn who goes on a quest to save the Herla, the deer kind.[114] In the 1942 Walt Disney Pictures film, Bambi a oq dumli kiyik, ichida Feliks Salten 's original 1923 book Bambi, a Life in the Woods, u a kiyik. Yilda C. S. Lyuis 's 1950 fantasy novel Arslon, jodugar va shkaf the adult Pevensies, now kings and queens of Narniya, chase the White Stag on a hunt, as the Stag is said to grant its captor a wish. The hunt is key in returning the Pevensies to their home in England. In the 1979 book Farting yog'ochining hayvonlari, The Great White Stag is the leader of all the animals.

Heraldiya

Qurollari Raon-o-Bois, Frantsiya

Deer of various types appear frequently in European geraldika. In the British armory, the term "stag" is typically used to refer to antlered male red deer, while "buck" indicates an antlered male fallow deer. Stags and bucks appear in a number of munosabat, referred to as "lodged" when the deer is lying down, "trippant" when it has one leg raised, "courant" when it is running, "springing" when in the act of leaping, "statant" when it is standing with all hooves on the ground and looking ahead, and "at gaze" when otherwise statant but looking at the viewer. Stags' heads are also frequently used; these are typically portrayed without an attached neck and as facing the viewer, in which case they are termed "caboshed".[115]

Examples of deer in gerblar can be found in the arms of Xertfordshir, England, and its county town of Xertford; both are examples of qo'llar. A deer appears on the arms of the Israeli Postal Authority. Coats of arms featuring deer include those of Dotternhauzen, Thierachern, Frioltsgeym, Bauen, Albstadt va Dassel Germaniyada; ning Earls Bathurst Angliyada; ning Balaxna, Rossiya; ning Alandiya, Finland; ning Gjemnes, Xitra, Xyartdal, Rendalen va Voss in Norway; ning Jeleniya Gora, Polsha; ning Umea, Shvetsiya; ning Kvinslend, Avstraliya; ning Cervera, Catalonia; ning Shimoliy Irlandiya; va Chili.[iqtibos kerak ]

Other types of deer used in heraldry include the hind, portrayed much like the stag or buck but without antlers, as well as the reindeer and winged stags. Winged stags are used as tarafdorlari in the arms of the de Carteret family. The sea-stag, possessing the antlers, head, forelegs and upper body of a stag and the tail of a suv parisi, is often found in German heraldry.[115]

Iqtisodiy ahamiyati

Deer have long had economic significance to humans. Deer meat, known as kiyik go'shti, is highly nutritious.[116][117] Due to the inherently wild nature and diet of deer, venison is most often obtained through deer hunting. In the United States, it is produced in small amounts compared to beef but still represents a significant trade. Conservation laws prevent the sale of wild game meat, although it may be donated. By 2012, some 25,000 tons of red deer were raised on farms in North America. The major deer-producing countries are New Zealand, the market leader, with Ireland, Great Britain and Germany. The trade earns over $100 million annually for these countries.[118]

The skins make a peculiarly strong, soft leather, known as buckskin. There is nothing special about skins with the fur on since the hair is brittle and soon falls off. The hoofs and horns are used for ornamental purposes, especially the antlers of the kiyik, which are utilized for making umbrella handles, and for similar purposes; elk horn is often employed in making knife handles. In China, a medicine is made from stag horn, and the antlers of certain species are eaten when "in the velvet".[119] Orasida Inuit, an'anaviy ulu women's knife was made with an antler, horn, or ivory handle.[120]

Nicholas Mavrogenes, Fanariot Shahzoda ning Valaxiya, riding through Buxarest in a stag−drawn carriage. Late 1780s

Deer have long been bred in captivity as ornaments for parks, but only in the case of reindeer has thorough domestication succeeded.[119] The Sami of Scandinavia and the Kola yarim oroli of Russia and other nomadic peoples of northern Asia use reindeer for food, clothing, and transport. Deer bred for hunting are selected based on the size of the antlers.[121] In North America, the reindeer, known there as caribou, is not domesticated or herded, but it is important as a quarry animal to the Caribou Inuit.[122]

Automobile collisions with deer can impose a significant cost on the economy. In the U.S., about 1.5 million deer-vehicle collisions occur each year, according to the National Highway Traffic Safety Administration. Those accidents cause about 150 human deaths and $1.1 billion in property damage annually.[123] In Scotland, several roads including the A82, A87 va A835 have had significant enough problems with deer vehicle collisions (DVCs) that sets of vehicle activated automatic warning signs have been installed along these roads.[124]

In some areas of the UK, deer (especially quruq kiyik due to their gregarious behaviour ), have been implicated as a possible reservoir for transmission of sigirning sil kasalligi,[46][47] a disease which in the UK in 2005 cost £90 million in attempts to eradicate.[48] In New Zealand, deer are thought to be important as vectors picking up M. bovis in areas where brushtail possums Trichosurus vulpecula are infected, and transferring it to previously uninfected possums when their carcasses are scavenged elsewhere.[49] The white-tailed deer Odocoileus virginianus has been confirmed as the sole maintenance host in the Michigan outbreak of bovine tuberculosis which remains a significant barrier to the US nationwide eradication of the disease in livestock. In 2008, 733,998 licensed deer hunters killed approximately 489,922 white-tailed deer to procure venison, control the deer population, and minimize the spread of disease. These hunters purchased more than 1.5 million deer harvest tags. The economic value of deer hunting to Michigan's economy is substantial. For example, in 2006, hunters spent US$507 million hunting white-tailed deer in Michigan.[50]

Deer hunting is a popular activity in the U.S. that provides the hunter's family with high quality meat and generates revenue for states and the federal government from the sales of licenses, permits and tags. The 2006 survey by the AQSh baliq va yovvoyi tabiatni muhofaza qilish xizmati estimates that license sales generate approximately $700 million annually. This revenue generally goes to support conservation efforts in the states where the licenses are purchased. Overall, the U.S. Fish and Wildlife Service estimates that big game hunting for deer and elk generates approximately $11.8 billion annually in hunting-related travel, equipment and related expenditures.[125]

Etimologiya

So'z kiyik was originally broad in meaning, becoming more specific with time. Qadimgi ingliz dēor va O'rta ingliz der meant a wild animal of any kind. Cognates of Old English dēor in other dead German tillari have the general sense of hayvon, kabi Qadimgi yuqori nemis tior, Qadimgi Norse djur yoki dȳr, Gotik dius, Qadimgi Sakson dierva Qadimgi friz diar.[126] This general sense gave way to the modern English sense by the end of the Middle English period, around 1500. However, all modern Germanic languages save English and Scots retain the more general sense: for example, German Qatlam and Norwegian dyr anglatadi hayvon.[127]

Terminologiya

"The Stag Hunt of Frederik III, Saksoniya saylovchisi "tomonidan Katta Lukas Kranax, 1529

For many types of deer in modern English usage, the male is a buk and the female a doe, but the terms vary with dialect, and according to the size of the species. Erkak qizil kiyik a qoqmoq, while for other large species the male is a buqa, the female a sigir, as in cattle. In older usage, the male of any species is a xart, especially if over five years old, and the female is a hind, especially if three or more years old.[128] The young of small species is a qirg'iy and of large species a calf; a very small young may be a bola. A castrated male is a havier.[129] A group of any species is a podani. The sifat of relation is cervine; like the family name Cervidae, this is from Lotin: cervus, ma'no qoqmoq yoki kiyik.

Shuningdek qarang

Adabiyotlar

  1. ^ 'deer' singular and plural among examples (swine OE swin, deer OE deor, sheep OE sceap, horse OE hors, year OE gear, pound OE pana) -Jespersen, A Modern English Grammar on Historical Principles, Part II SYNTAX (First Volume), Ch.III The Unchanged Plural (p. 49) arrow.latrobe.edu.au accessed 14 November 2020
  2. ^ Iltanen, Jussi: Suomen kuntavaakunat (2013), Karttakeskus, ISBN  951-593-915-1
  3. ^ McLintock, A. H., ed. (1966). "Deer". Te Ara: An Encyclopaedia of New Zealand.
  4. ^ a b Groves, C. (2007). "Family Cervidae". Yilda Prothero, D. R.; Foss, S. E. (eds.). The Evolution of Artiodactyls (Tasvirlangan tahrir). Baltimore, USA: Johns Hopkins University Press. pp. 249–56. ISBN  978-0-801-88735-2.
  5. ^ Kingdon, J. (2015). The Kingdon Field Guide to African Mammals (2-nashr). London, UK: Bloomsbury Publishing. p. 499. ISBN  978-1-4729-2531-2.
  6. ^ Jameson, E. W.; Peeters, H. J., Jr. (2004). Mammals of California (Qayta ko'rib chiqilgan tahrir). Berkeley, USA: University of California Press. p. 241. ISBN  978-0-520-23582-3.
  7. ^ Long, C. A. (2008). The Wild Mammals of Wisconsin. Sofia, Bulgaria: Pensoft. p.439. ISBN  9789546423139.
  8. ^ Prothero, D. R.; Schoch, R. M. (2002). Horns, Tusks, and Flippers: The Evolution of Hoofed Mammals. Baltimore, USA: Johns Hopkins University Press. pp. 61–84. ISBN  978-0-8018-7135-1.
  9. ^ a b Kurta, A. (1995). Mammals of the Great Lakes Region (1-nashr). Michigan, USA: University of Michigan Press. pp.260–1. ISBN  978-0-472-06497-7.
  10. ^ a b v d e f g Geist, V. (1998). Deer of the World: Their Evolution, Behaviour and Ecology (1-nashr). Mechanicsburg, USA: Stackpole Books. pp. 1–54. ISBN  978-0-8117-0496-0.
  11. ^ Armstrong, D. M.; Fitzgerald, J. P.; Meaney, C. A. (2011). Mammals of Colorado (2-nashr). Colorado, USA: University Press of Colorado. p. 445. ISBN  978-1-60732-048-7.
  12. ^ Kingdon, J.; Xappold, D .; Butinski, T .; Hoffmann, M.; Xappold, M .; Kalina, J. (2013). Afrikaning sutemizuvchilar. VI. London, UK: Bloomsbury Publishing. p. 116. ISBN  978-1-4081-8996-2.
  13. ^ a b Feldhamer, G. A.; McShea, W. J. (2012). Deer: The Animal Answer Guide. Baltimore, USA: Johns Hopkins University Press. pp. 1–142. ISBN  978-1-4214-0387-8.
  14. ^ Francis, C. M. (2008). A Field Guide to the Mammals of South-East Asia. London, UK: New Holland. p. 130. ISBN  978-1-84537-735-9.
  15. ^ Trolle, M.; Emmons, L. H. (2004). "A record of a dwarf brocket from Lowland Madre De Dios, Peru" (PDF). Deer Specialist Group News (19): 2–5.
  16. ^ Schmidly, D. J. (2004). The Mammals of Texas (Qayta ko'rib chiqilgan tahrir). Austin, Texas (USA): University of Texas Press. 263-4 betlar. ISBN  978-1-4773-0886-8.
  17. ^ Hames, D. S.; Koshowski, Denise (1999). Hoofed Mammals of British Columbia. Vancouver, Canada: UBC Press. p. 113. ISBN  978-0-7748-0728-9.
  18. ^ Booy, O.; Wade, M.; Roy, H. (2015). Field Guide to Invasive Plants and Animals in Britain. London, UK: Bloomsbury Publishing. p. 170. ISBN  978-1-4729-1153-7.
  19. ^ Bowers, N.; Bowers, R.; Kaufmann, K. (2004). Mammals of North America. New York, USA: Houghton Mifflin. pp.158–9. ISBN  978-0-618-15313-8.
  20. ^ Hooey, T. (2004). Strategic Whitetail Hunting. Krause nashrlari. p. 39. ISBN  978-1-4402-2702-8.
  21. ^ Ryder, M. L.; Kay, R. N. B. (1973). "Structure of and seasonal change in the coat of Red deer (Servus elafusi)". Zoologiya jurnali. 170 (1): 69–77. doi:10.1111/j.1469-7998.1973.tb05044.x.
  22. ^ Lincoln, G. A.; Guinness, F. E. (1972). "Effect of altered photoperiod on delayed implantation and moulting in roe deer" (PDF). Ko'paytirish. 31 (3): 455–7. doi:10.1530/jrf.0.0310455. PMID  4648129.
  23. ^ Owen, James (25 August 2003). "Scottish Deer Are Culprits in Bird Killings". National Geographic News. Olingan 16 iyun 2009.
  24. ^ Dale, Michael (1988). "Carnivorous Deer". Omni Magazine: 31.
  25. ^ Burton, M.; Burton, R. (2002). Xalqaro yovvoyi tabiat entsiklopediyasi (3-nashr). New York, US: Marshall Cavendish. pp.446–7. ISBN  978-0-7614-7270-4.
  26. ^ a b v d e Hall, B. K. (2005). Bones and Cartilage: Developmental and Evolutionary Skeletal Biology. Amsterdam, Netherlands: Elsevier Academic Press. pp. 103–15. ISBN  978-0-08-045415-3.
  27. ^ Wislocki, G. B. (1954). "Antlers in female deer, with a report of three cases in Odocoileus". Mammalogy jurnali. 35 (4): 486–95. doi:10.2307/1375571. JSTOR  1375571.
  28. ^ Smith, T. (2013). The Real Rudolph: A Natural History of the Reindeer. New York, US: The History Press. ISBN  978-0-7524-9592-7.
  29. ^ Fletcher, T. J. (1986). "Reproduction: seasonality". In Alexander, T. L.; Buxton, D. (eds.). Management and Diseases of Deer: A Handbook for the Veterinary Surgeon (2-nashr). London, UK: Veterinary Deer Society. pp. 17–8. ISBN  978-0-9510826-0-7.
  30. ^ Malo, A. F.; Roldan, E. R. S.; Garde, J.; Soler, A. J.; Gomendio, M. (2005). "Antlers honestly advertise sperm production and quality". Qirollik jamiyati materiallari B: Biologiya fanlari. 272 (1559): 149–57. doi:10.1098/rspb.2004.2933. PMC  1634960. PMID  15695205.
  31. ^ Emlen, D. J. (2008). "The evolution of animal weapons". Annual Review of Ecology, Evolution, and Systematics. 39: 387–413. doi:10.1146/annurev.ecolsys.39.110707.173502.
  32. ^ Bowyer, R. T. (1986). "Antler characteristics as related to social status of male southern mule deer". Janubi-g'arbiy tabiatshunos. 31 (3): 289–98. doi:10.2307/3671833. JSTOR  3671833.
  33. ^ Ditchkoff, S. S.; Lochmiller, R. L.; Masters, R. E.; Hoofer, S. R.; Den Bussche, R. A. Van (2001). "Major-histocompatibility-complex-associated variation in secondary sexual traits of white-tailed deer (Odocoileus virginianus) evidence for good-genes advertisement". Evolyutsiya. 55 (3): 616–625. doi:10.1111/j.0014-3820.2001.tb00794.x. PMID  11327168. S2CID  10418779.
  34. ^ Malo, A. F.; Roldan, E. R. S.; Garde, J.; Soler, A. J.; Gomendio, M. (2005). "Antlers honestly advertise sperm production and quality". Qirollik jamiyati materiallari B: Biologiya fanlari. 272 (1559): 149–157. doi:10.1098/rspb.2004.2933. PMC  1634960. PMID  15695205.
  35. ^ Metz, Matthew C.; Emlen, Douglas J.; Stahler, Daniel R.; MacNulty, Daniel R.; Smith, Douglas W. (3 September 2018). "Predation shapes the evolutionary traits of cervid weapons". Tabiat ekologiyasi va evolyutsiyasi. 2 (10): 1619–1625. doi:10.1038/s41559-018-0657-5. PMID  30177803. S2CID  52147419.
  36. ^ Garrod, A. Notes on the visceral anatomy and osteology of the ruminants, with a suggestion regarding a method of expressing the relations of species by means of formulae. Proceedings of the Zoological Society of London, 2–18 (1877).
  37. ^ Brooke, V. On the classification of the Cervidæ, with a synopsis of the existing Species. Journal of Zoology 46, 883–928 (1878).
  38. ^ Pocock, R. The Homologies between the Branches of the Antlers of the Cervidae based on the Theory of Dichotomous Growth. Journal of Zoology 103, 377–406 (1933).
  39. ^ Samejima, Y., Matsuoka, H. A new viewpoint on antlers reveals the evolutionary history of deer (Cervidae, Mammalia). Sci Rep 10, 8910 (2020). https://doi.org/10.1038/s41598-020-64555-7
  40. ^ Reid, F. A. (2006). A Field Guide to Mammals of North America, North of Mexico (4-nashr). Boston, USA: Houghton Mifflin Co. pp. 153–4. ISBN  978-0-395-93596-5.
  41. ^ Cockerill, R. (1984). Makdonald, D. (tahrir). Sutemizuvchilar entsiklopediyasi. New York, USA: Facts on File. pp.520–9. ISBN  978-0-87196-871-5.
  42. ^ Uresk, Daniel W., and Donald R. Dietz. "Fecal vs. Rumen Contents to Determine White-tailed Deer Diets." Intermountain Journal of Sciences 24, no. 3-4 (2018): 118-122.
  43. ^ Case, D.J.; McCullough, D.R. (February 1987). "White-tailed deer forage on alewives". Mammalogy jurnali. 68 (1): 195–198. doi:10.2307/1381075. JSTOR  1381075.
  44. ^ Ellis-Felege, S. N.; Burnam, J. S.; Palmer, W. E.; Sisson, D. C.; Wellendorf, S. D.; Thornton, R. P.; Stribling, H. L.; Carroll, J. P. (2008). "Cameras identify White-tailed deer depredating Northern bobwhite nests". Southeastern Naturalist. 7 (3): 562–564. doi:10.1656/1528-7092-7.3.562. S2CID  84790827.
  45. ^ Deer – info and games Sheppard Software.
  46. ^ a b Delahay, R. J.; Smith, G. C.; Barlow, A. M.; Walker, N.; Harris, A.; Clifton-Hadley, R. S.; Cheeseman, C. L. (2007). "Bovine tuberculosis infection in wild mammals in the South-West region of England: A survey of prevalence and a semi-quantitative assessment of the relative risks to cattle". The Veterinary Journal. 173 (2): 287–301. doi:10.1016/j.tvjl.2005.11.011. PMID  16434219.
  47. ^ a b Ward, A. I.; Smith, G. C.; Etherington, T. R.; Delahay, R. J. (2009). "Estimating the risk of cattle exposure to tuberculosis posed by wild deer relative to badgers in England and Wales". Yovvoyi tabiat kasalliklari jurnali. 45 (4): 1104–1120. doi:10.7589/0090-3558-45.4.1104. PMID  19901384. S2CID  7102058.
  48. ^ a b Anonymous (2008). "Bovine TB: EFRACom calls for a multifaceted approach using all available methods". Veterinariya qaydlari. 162 (9): 258–259. doi:10.1136/vr.162.9.258. PMID  18350673. S2CID  2429198.
  49. ^ a b Delahay, R. J.; De Leeuw, A. N. S.; Barlow, A. M.; Clifton-Hadley, R. S.; Cheeseman, C. L. (2002). "The status of Mycobacterium bovis infection in UK wild mammals: A review". The Veterinary Journal. 164 (2): 90–105. doi:10.1053/tvjl.2001.0667. PMID  12359464.
  50. ^ a b O'Brien, D. J.; Schmitt, S. M.; Fitzgerald, S. D.; Berry, D. E. (2011). "Management of bovine tuberculosis in Michigan wildlife: Current status and near term prospects". Veterinariya mikrobiologiyasi. 151 (1–2): 179–187. doi:10.1016/j.vetmic.2011.02.042. PMID  21414734.
  51. ^ a b Alan Cochrane (January 2019). "Don't fraternize with wild animals: biologist". Moncton Times&Transcript.
  52. ^ "Wildlife and Heritage Service : Chronic Wasting Disease (CWD)". Merilend tabiiy resurslar departamenti. Arxivlandi asl nusxasi 2013 yil 14 mayda.
  53. ^ a b v d Goss, R. J. (1983). Deer Antlers Regeneration, Function and Evolution. Oxford, UK: Elsevier. pp. 43–51. ISBN  9780323140430.
  54. ^ a b Janis, C. M .; Effinger, J. A.; Harrison, J. A.; Honey, J. G.; Kron, D. G.; Lander, B.; Manning, E.; Prothero, D.; Stevens, M. S.; Stucky, R. K.; Webb, S. D.; Wright, D. B. (1998). "Artiodactyla". In Janis, C. M.; Scott, K. M.; Jacobs, L. L. (eds.). Evolution of Tertiary Mammals of North America (1-nashr). Kembrij, Buyuk Britaniya: Kembrij universiteti matbuoti. pp.337 –74. ISBN  978-0-521-35519-3.
  55. ^ a b Heffelfinger, J. (2006). Deer of the Southwest : A Complete Guide to the Natural History, Biology, and Management of Southwestern Mule Deer and White-tailed Deer (1-nashr). Texas, USA: Texas A & M University Press. 1-57 betlar. ISBN  978-1-58544-515-8.
  56. ^ Rose, K. D. (1982). "Skeleton of Diacodexis, oldest known artiodactyl". Ilm-fan. 216 (4546): 621–3. Bibcode:1982Sci...216..621R. doi:10.1126/science.216.4546.621. JSTOR  1687682. PMID  17783306. S2CID  13157519.
  57. ^ Eldredge, N.; Stanley, S. M., eds. (1984). Tirik qoldiqlar. New York, USA: Springer. ISBN  978-1-4613-8271-3.
  58. ^ a b v Ludt, C. J.; Schroeder, W.; Rottmann, O.; Kuehn, R. (2004). "Mitochondrial DNA phylogeography of red deer (Servus elafusi)". Molekulyar filogenetik va evolyutsiyasi. 31 (3): 1064–83. doi:10.1016/j.ympev.2003.10.003. PMID  15120401.
  59. ^ Vislobokova, I.; Daxner-Höck, G. (2001). "Oligocene–early Miocene ruminants from the Valley of Lakes (central Mongolia)" (PDF). Annalen des Naturhistorischen Museums in Wien. A. 103: 213–35. JSTOR  41702231.
  60. ^ Stirton, R. A. (1944). "Comments on the relationships of the cervoid family Palaeomerycidae". Amerika Ilmiy jurnali. 242 (12): 633–55. Bibcode:1944AmJS..242..633S. doi:10.2475/ajs.242.12.633.
  61. ^ Agate Fossil Beds: Agate Fossil Beds National Monument, Nebraska. Interior Department, National Park Service, Division of Publications. February 1989. p. 31. ISBN  978-0-912627-04-5.
  62. ^ Gentry, A. W.; Rössner, G. (1994). "The Miocene differentiation of Old World Pecora (Mammalia)". Tarixiy biologiya. 7 (2): 115–58. doi:10.1080/10292389409380449.
  63. ^ Azanza, B.; DeMiguel, D.; Andrés, M. (2011). "The antler-like appendages of the primitive deer Dicrocerus elegans: morphology, growth cycle, ontogeny, and sexual dimorphism". Estudios Geológicos. 67 (2): 579–602. doi:10.3989/egeol.40559.207.
  64. ^ Vang X.; Xie, G.; Dong, W. (2009). "Toj shoxli kiyikning yangi turi Stefanokemalar (Artiodactyla, Cervidae) Qaydam havzasining o'rta miosenidan, Shimoliy Tibet platosi, Xitoy va uning filogeniyasini oldindan baholash ". Linnean Jamiyatining Zoologik jurnali. 156 (3): 680–95. doi:10.1111 / j.1096-3642.2008.00491.x.
  65. ^ Ginsburg, L. (1988). "La faune des mammifères des sables Miocènes du synclinal d'Esvres (Val de Loire)" [Esvres (Luara vodiysi) sinxlinasining miosen qumlarining sutemizuvchilar faunasi]). Comptes Rendus de l'Académie des Sciences. II (frantsuz tilida): 319–22.
  66. ^ Walker, D. N. (2000). "Pleystotsen va golotsen yozuvlari Antilocapra americana: FAUNMAP ma'lumotlarini ko'rib chiqish ". (PDF). Tekislik antropologi. 45 (174): 13–28. doi:10.1080/2052546.2000.11932020. JSTOR  25669684. S2CID  163903264.
  67. ^ Dong, V.; Pan, Y .; Liu, J. (2004 yil sentyabr). "Eng qadimgi Muntiakus (Artiodactyla, sutemizuvchilar) Yuanmou kech miosenidan, Xitoyning janubi-g'arbiy qismida ". Comptes Rendus Palevol. 3 (5): 379–86. doi:10.1016 / j.crpv.2004.06.002.
  68. ^ a b v d Gilbert, C .; Ropiquet, A .; Hassanin, A. (2006). "Servidlarning mitoxondriyal va yadro filogeniyalari (sutemizuvchilar, Ruminantiya): sistematika, morfologiya va biogeografiya". Molekulyar filogenetik va evolyutsiyasi. 40 (1): 101–17. doi:10.1016 / j.ympev.2006.02.017. PMID  16584894.
  69. ^ Di Stefano, G.; Petronio, C. (2002). "Evroosiyo plio-pleystotsen qabilasi Tservini sistematikasi va evolyutsiyasi (Artiodactyla, Mammalia)" (PDF). Geologica Romana. 36: 311-34. Arxivlandi asl nusxasi (PDF) 2016 yil 10 martda. Olingan 11 aprel 2016.
  70. ^ Petronio, C .; Kraxmalnaya, T .; Belluchchi, L .; Di Stefano, G. (2007). "Ba'zi bir Evroosiyo pliotservinlari haqida eslatmalar: xususiyatlari, evolyutsiyasi va Tservini qabilasi bilan aloqalari". Geobios. 40 (1): 113–30. doi:10.1016 / j.geobios.2006.01.002.
  71. ^ G'affor, A .; Axtar, M .; Nayyer, A. Q. (2011). "Pokiovaning Sivaliklaridan Tservini (Mammalia, Artiodactyla, Cervidae) qabilasining dastlabki pliyotsen qoldiqlari qoldiqlari to'g'risida dalillar" (PDF). Hayvonot va o'simlik fanlari jurnali. 21 (4): 830–5.
  72. ^ van der Made, J .; Morales, J .; Sen, S .; Aslan, F. (2002). "Turkiyaning yuqori miosen davridagi birinchi tuya va tuyalarning eski dunyoga tarqalishi". Comptes Rendus Palevol. 1 (2): 117–22. doi:10.1016 / S1631-0683 (02) 00012-X.
  73. ^ Vebb, S. D. (2000). "Yangi Dunyo Tservidasining evolyutsion tarixi". Yilda Vrba, E. S.; Shaller, G. B. (tahr.). Antilopalar, kiyiklar va qarindoshlar: fotoalbom yozuvlari, xulq-atvori ekologiyasi, sistematikasi va muhofazasi. Nyu-Xeyven, AQSh: Yel universiteti matbuoti. 38-64 betlar. ISBN  978-0-300-08142-8.
  74. ^ De Vos, J .; Mol, D .; Reumer, J. W. F. (1995). "Servisteusning revizyoni bilan Oosterscheldadan (Niderlandiya) erta pleystotsenli Servidae (Mammalia, Artiodactyla). Evkladokeros Falconer, 1868 " (PDF). "Deynsi" (2): 95–121.
  75. ^ Lister, A. M .; Gonsales, S .; Kitchener, A. C. (2000). "Irlandiyalik elkaning Golotsenga omon qolishi". Tabiat. 405 (6788): 753–4. Bibcode:2000. Natur.405..753G. doi:10.1038/35015668. PMID  10866185. S2CID  4417046.
  76. ^ Moen, R. A .; Pastor, J .; Yosef, C. (1999). "Irlandiyalik elkaning shox o'sishi va yo'q bo'lib ketishi" (PDF). Evolyutsion ekologiya tadqiqotlari (1): 235–49.
  77. ^ Breda, M .; Marchetti, M. (2005). "Plio-Pleistosen Altseyni (Cervidae; Mammalia) ning Evroosiyodan tizimli va bioxronologik tekshiruvi" (PDF). To'rtlamchi davrga oid ilmiy sharhlar. 24 (5–6): 775–805. Bibcode:2005QSRv ... 24..775B. doi:10.1016 / j.quascirev.2004.05.005.
  78. ^ a b Uilson, D.E.; Rider, D.M., tahr. (2005). Dunyoning sutemizuvchilar turlari: taksonomik va geografik ma'lumot (3-nashr). Jons Xopkins universiteti matbuoti. p. 652-70. ISBN  978-0-8018-8221-0. OCLC  62265494.
  79. ^ Bruk, V. (1878). "Cervidœ klassifikatsiyasi to'g'risida, mavjud turlarning sinopsi bilan". London zoologik jamiyati materiallari. 46 (1): 883–928. doi:10.1111 / j.1469-7998.1878.tb08033.x.
  80. ^ Groves, S; Grubb, P. (1987). "Tirik kiyikning munosabatlari". Vemmerda C. (tahrir). Cervidae biologiyasi va boshqaruvi: 1982 yil 1–5 avgust kunlari Virjiniya shtatidagi Front Royal, Smitson instituti, Milliy Zoologik Park, Tabiatni muhofaza qilish va tadqiqot markazida bo'lib o'tgan konferentsiya.. Vashington, AQSh: Smithsonian Institution Press. 21-59 betlar. ISBN  978-0-87474-981-6.
  81. ^ Grubb, P. (2000). "Tirik va yaroqli kiyiklarning haqiqiy va yaroqsiz nomenklaturasi, Cervidae" (PDF). Acta Theriologica. 45 (3): 289–307. doi:10.4098 / soat.arch.00-30.
  82. ^ a b Xasanin, A .; Douzery, E. J. P. (2003). "Ruminantiyaning molekulyar va morfologik filogeniyalari va Moschidae ning muqobil holati". Tizimli biologiya. 52 (2): 206–28. doi:10.1080/10635150390192726. PMID  12746147.
  83. ^ Randi, E .; Mucci, N .; Pierpaoli, M .; Douzery, E. (1998). "Cervidae (Artiodactyla) ning yangi filogenetik istiqbollari mitoxondriyal sitoxrom b geni tomonidan ta'minlanadi". Qirollik jamiyati materiallari B: Biologiya fanlari. 265 (1398): 793–801. doi:10.1098 / rspb.1998.0362. PMC  1689037. PMID  9628037.
  84. ^ Randi, E .; Mucci, N .; Klaro-Xergueta, F.; Kapot, A .; Douzery, E. J. P. (2001). "Serviksning mitoxondriyal DNK nazorati mintaqasi filogeniyasi: spetsifikatsiya Servus va tabiatni muhofaza qilishning oqibatlari ". Hayvonlarni muhofaza qilish. 4 (1): 1–11. doi:10.1017 / S1367943001001019.
  85. ^ Lyudt, C. J .; Shreder, V.; Rottmann, O .; Kuehn, R. (2004). "Qizil kiyikning mitoxondrial DNK filogeografiyasi (Servus elafusi)". Molekulyar filogenetik va evolyutsiyasi. 31 (3): 1064–83. doi:10.1016 / j.ympev.2003.10.003. PMID  15120401.
  86. ^ Fernández, M. H.; Vrba, E. S. (2005). "Ruminantiyadagi filogenetik munosabatlarning to'liq bahosi: mavjud bo'lgan kavsh qaytaruvchi hayvonlarning turlar darajasidagi supertree". Biologik sharhlar. 80 (2): 269–302. doi:10.1017 / S1464793104006670. PMID  15921052. S2CID  29939520.
  87. ^ Ruis-Garsiya, M.; Randi, E .; Martines-Agüero, M.; Alvarez, D. (iyun 2007). "Netropik kiyik avlodlari (Artiodactyla: Cervidae) o'rtasida mtDNA ketma-ketliklari va mikrosatellit markerlari orqali filogenetik munosabatlar". Revista de Biología Tropical. 55 (2): 723–41. PMID  19069784.
  88. ^ Duarte, J. M. B.; Gonsales, S .; Maldonado, J. E. (2008). "Janubiy Amerika kiyiklarining ajablanarli evolyutsion tarixi". Molekulyar filogenetik va evolyutsiyasi. 49 (1): 17–22. doi:10.1016 / j.ympev.2008.07.009. PMID  18675919.
  89. ^ Pitra, C .; Fikel, J .; Meyxard, E .; Groves, S (2004). "Qadimgi dunyo kiyiklari evolyutsiyasi va filogeniyasi". Molekulyar filogenetik va evolyutsiyasi. 33 (3): 880–895. doi:10.1016 / j.ympev.2004.07.013. PMID  15522810.
  90. ^ Gentri, A. V. (1993). "Procervulus Gaudri, 1877 yil (Mammalia, Artiodactyla): tavsiya etilgan nom Antilopik dixotoma Gervais, 1849 yil tur sifatida ". Zoologik nomenklatura byulleteni. 50 (2): 137–9. doi:10.5962 / bhl.part.1820.
  91. ^ Dong, V.; Liu, J .; Pan, Y. (2003). "Yangi Evropoks Xitoyning Yunnan viloyati, Yuanmou shahridagi so'nggi miosen, uning atrof muhitini sharhlagan holda ". Xitoy fanlari byulleteni. 48 (5): 485–91. Bibcode:2003ChSBu..48..485D. doi:10.1007 / BF03183257. S2CID  140586480.
  92. ^ Deng, T. (2014). "Ning yangi turi Eostilotseros (Cervidae, Artiodactyla) Xitoyning Gansu shahridagi Linxiya havzasining so'nggi miosenidan ". Zootaxa. 3893 (3): 363–81. doi:10.11646 / zootaxa.3893.3.3. PMID  25544527.
  93. ^ "Paracervulus". Turlar. GBIF. Olingan 14 aprel 2016.
  94. ^ Vang X.; Xie, G.; Dong, W. (2009). "Toj shoxli kiyikning yangi turi Stefanokemalar (Artiodactyla, Cervidae) Qaydam havzasining o'rta miosenidan, Shimoliy Tibet platosi, Xitoy va uning filogeniyasini oldindan baholash ". Linnean Jamiyatining Zoologik jurnali. 156 (3): 680–95. doi:10.1111 / j.1096-3642.2008.00491.x.
  95. ^ Petronio, C .; Pandolfi, L. (2011). "Arvernoceros Heintz jinsining birinchi marta paydo bo'lishi, 1970 yil Italiyaning pleystotsen davrining oxirlarida". Rivista Italiana di Paleontologia e Stratigrafia. 117 (3): 501–8. doi:10.13130/2039-4942/5988.
  96. ^ Lindsay, E. H.; Falbuss, V .; Mein, P., nashr. (1989). Evropa neogen sutemizuvchilar xronologiyasi. Nyu-York, AQSh: Plenum matbuoti. p. 205. ISBN  978-1-4899-2513-8.
  97. ^ Amerika Qo'shma Shtatlari Geologiya xizmati byulleteni (Hisobot). Amerika Qo'shma Shtatlarining Geologik xizmati. 1902. 682-6 betlar.
  98. ^ Vislobokova, I.A. (2011 yil dekabr). "Gigant kiyiklarning tarixiy rivojlanishi va geografik tarqalishi (Cervidae, Megacerini)". Paleontologik jurnal. 45 (6): 674–688. doi:10.1134 / S0031030111060153. S2CID  85326341.
  99. ^ "Pliyotservus". Turlar. GBIF. Olingan 14 aprel 2016.
  100. ^ Mazza, P. P. A. (2006). "Poggio Rosso (Yuqori Valdarno, markaziy Italiya), so'nggi Pliyotsen yovvoyi hayoti haqidagi oyna". PALAY. 21 (5): 493–8. Bibcode:2006 yil Palay..21..493M. doi:10.2110 / palo.2005.P05-033R. S2CID  129513994.
  101. ^ Morejon, G.V .; Xerst, J .; va boshq. (2005 yil fevral). "Yo'qolib ketgan serviks turini postkranial karpalial qo'llab-quvvatlash Bretziya bog'liq shox bilan ". Mammalogy jurnali. 86 (1): 115–120. doi:10.1644 / 1545-1542 (2005) 086 <0115: PCSFTE> 2.0.CO; 2.
  102. ^ Valli, A.M.F. (2010 yil fevral). "Jinsning tarqalishi Prokapreol va o'rtasidagi munosabatlar Prokapreolus cusanus va kiyik (Kapreol)". To'rtlamchi xalqaro. 212 (2): 80–85. Bibcode:2010QuInt.212 ... 80V. doi:10.1016 / j.quaint.2008.11.002.
  103. ^ Flerov, K.K .; Shevyreva, N.S. (1965). "Pseudalces, Kiskavkazdan kelgan pliosen kiyiklari ". Xalqaro geologiya sharhi. 7 (5): 931–934. Bibcode:1965IGRv .... 7..931F. doi:10.1080/00206816509474162.
  104. ^ Valli, A.M.F. (Mart 2018). "Serra da Capivara" (Piauí, Braziliya) milliy bog'i hududida pleystotsenning so'nggi ko'ruvchisi ". To‘rtlamchi davr. 1 (1): 4. doi:10.3390 / quat1010004.
  105. ^ Uebb, S.D .; Xulbert, RC; va boshq. (2008 yil may). "Markaziy Florida Fosfat okrugidan Palmetto faunasining quruqlikdagi sutemizuvchilari (pliyosenning boshi, eng so'nggi Hemphillian)". Ilmiy seriyalar, Los-Anjeles okrugining tabiiy tarix muzeyi. 41: 293–312. Olingan 12 avgust 2020.
  106. ^ Alkaraz, M .; Zurita, A. (2004). "Nuevos registros de cérvidos poco conocidos: Epieuryceros qarang proximus Castellanos y Antifer sp. (Sutemizuvchilar, Artiodactyla, Cervidae) ". Revista de Museo Argentino de Ciencias Naturales. 6 (1). Olingan 12 avgust 2020.
  107. ^ a b v d Feldhamer, G. A .; McShea, W. J. (2011). Kiyik: Hayvonlarga javob berish bo'yicha qo'llanma. Baltimor: Jons Xopkins universiteti matbuoti. 123-32 betlar. ISBN  9781421403885.
  108. ^ Kurtis, Gregori (2006). G'or rassomlari: dunyodagi birinchi rassomlarning sirlarini tekshirish (1-nashr). Nyu-York: Alfred A. Knopf. pp.96–97, 102. ISBN  978-1400043484.
  109. ^ Chugg, Endryu (2006). Aleksandrning sevgililari. Raleigh, N.C .: Lulu. 78-79 betlar. ISBN  978-1-4116-9960-1.
  110. ^ Harding, Endryu (2006 yil 23 sentyabr). "Pekin jinsiy olatni emporium". BBC yangiliklari. Olingan 23 iyun 2010.
  111. ^ Berrin, Ketrin va Larko muzeyi (1997) Qadimgi Peru ruhi: dan xazinalar Museo Arqueológico Rafael Larco Herrera. Nyu York: Temza va Xadson, ISBN  0500018022.
  112. ^ Xarvi, G. (2002). Mahalliy dinlarda o'qishlar. London: doimiylik. p. 109. ISBN  978-0826451019.
  113. ^ Mur, Klement S (1823 yil 2-dekabr). "Aziz Nikolayning tashrifi haqida hisobot". Troy Sentinel. p. 2018-04-02 121 2. Olingan 27 mart 2015.
  114. ^ Klement-Devis, D. (2007). Fire Bringer (1-Amerika nashri). Nyu-York: Firebird. ISBN  978-0142408735.
  115. ^ a b Artur Foks-Devis, Heraldiya uchun to'liq qo'llanma, T.C. va E.C. Jek, London, 1909, 208-210,
  116. ^ Kralj, Richard Endryu (sentyabr 2014). "Venison, bu siz uchunmi?". Penn State kengaytmasi. Olingan 20 yanvar 2020.
  117. ^ Blitman, Joanna; Sykes, Rozi (2013 yil sentyabr). "Nima uchun kiyik go'shti sizga foydalidir | Joanna Blytman va Rozi Syks". The Guardian. Olingan 20 yanvar 2020.
  118. ^ Burden, Dan (iyun 2012). "Deer Venison Ranching profili". Qishloq xo'jaligi marketingining resurs markazi. Olingan 11 aprel 2016.
  119. ^ a b Rines, Jorj Edvin, ed. (1920). "Kiyik". Entsiklopediya Amerika.
  120. ^ "Inuit Bering Sea Eskimo Walrus fil suyagi va temir yarim oy pichog'i" Ulu "(1800 dan 1900 gacha Inuit)". Olingan 2 oktyabr 2018.
  121. ^ Laskow, Sara (2014 yil 27-avgust). "Bug'doy fermasi". O'rta (xizmat). Olingan 28 avgust 2014.
  122. ^ "Karibu Inuit". everyculture.com. Olingan 11 aprel 2016.
  123. ^ "Bug'u avtohalokatlari uchun eng yomon holatlar". CNN.com. 2006 yil 14-noyabr. Olingan 5 aprel 2009.
  124. ^ "Shimoliy G'arbiy hudud: Avtomobilda faollashtirilgan kiyikni ogohlantirish belgilari" (PDF). Transport Shotlandiya. Aprel 2010. 07 / NW / 0805/046. Olingan 11 iyul 2013. Iqtibos jurnali talab qiladi | jurnal = (Yordam bering)
  125. ^ "AQSh Ichki ishlar vazirligi, Baliq va yovvoyi tabiatni muhofaza qilish vazirligi va AQSh Savdo vazirligi, AQSh aholini ro'yxatga olish byurosi. 2006 yil baliq ovlash, ov qilish va yovvoyi tabiat bilan bog'liq bo'lgan milliy tadqiqotlar" (PDF). Olingan 16 noyabr 2012.
  126. ^ "kiyik". Ingliz tilining Amerika merosi lug'ati (4-nashr). Houghton Mifflin kompaniyasi. 2000. Arxivlangan asl nusxasi 2004 yil 25 martda.
  127. ^ Xarper, Duglas. "Kiyik". Onlayn etimologiya lug'ati. Olingan 7 iyun 2012.
  128. ^ OED, s.v. xart va orqada
  129. ^ "Havier". Dictionary.com. Olingan 4 avgust 2012.

Qo'shimcha o'qish

  • Deerland: Amerikaning ekologik muvozanat va yovvoyi tabiatning ovi Al Cambronne tomonidan, Lyons Press (2013), ISBN  978-0-7627-8027-3

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