Oddiy starling - Common starling

Oddiy starling
Oddiy starling
Oddiy starlingning qo'shig'i
Ilmiy tasnif tahrirlash
Qirollik:Animalia
Filum:Chordata
Sinf:Aves
Buyurtma:Passeriformes
Oila:Sturnidae
Tur:Sturnus
Turlar:
S. vulgaris
Binomial ism
Sturnus vulgaris
Sturnus vulgaris map.png
Mahalliy:
  Yozgi mehmon
  Rezident
  Qishki mehmon
Tanishtirdi:
  Yozgi mehmon
  Rezident

The oddiy starling (Sturnus vulgaris) deb nomlanuvchi Evropalik yulduzcha Qo'shma Shtatlarda yoki shunchaki starling Britaniya orollarida o'rta bo'yli passerin qush yulduzli oilada, Sturnidae. Uzunligi taxminan 20 sm (8 dyuym) va yaltiroq qora rangga ega tuklar yilning ba'zi paytlarida oq rangga bo'yalgan metall nashrida bilan. Oyoqlari pushti, qishda qora va yozda sariq rang; yosh qushlarning kattalarnikiga qaraganda jigarrang tuklari bor. Bu shovqinli qush, ayniqsa ichida jamoat uylari musiqiy bo'lmagan, ammo xilma-xil qo'shiq bilan va boshqa kulgili holatlar. Uning sovg'asi taqlid adabiyotda, jumladan, qayd etilgan Mabinogion va asarlari Katta Pliniy va Uilyam Shekspir.

Oddiy starlingda taxminan 12 ta pastki turlari ochiq joylarda ko'paytirish yashash joylari uning tug'ilgan joyi bo'ylab oralig'i mo''tadil Evropada va bo'ylab Palearktika g'arbiy Mo'g'ulistonga va u bilan tanishtirildi Avstraliya, Yangi Zelandiya, Kanada, Qo'shma Shtatlar, Meksika, Argentina, Janubiy Afrika va Fidji.[2] Ushbu qush Evropaning g'arbiy va janubiy qismida va Osiyoning janubi-g'arbiy qismida, populyatsiyasi shimoli-sharqida joylashgan ko'chib o'tish qishda janub va g'arbiy naslchilik doirasi ichida, shuningdek janubga qarab Iberiya va Shimoliy Afrika. Oddiy starling tartibsizlikni quradi uya to'rt yoki beshta porloq, xira ko'k tuxum qo'yilgan tabiiy yoki sun'iy bo'shliqda. Ularni tug'ilishi uchun ikki hafta vaqt ketadi va bolalar uyasida yana uch hafta qoladi. Odatda har yili bir yoki ikkita naslchilik urinishlari mavjud. Ushbu tur keng qamrovli turlarni o'z ichiga olgan omnivor hisoblanadi umurtqasizlar, shuningdek urug'lar va mevalar. Uni turli sutemizuvchilar va yirtqich qushlar va tashqi va ichki parazitlar mezbonidir.

Ushbu turga xos bo'lgan yirik podalar umurtqasiz hayvonlar zararkunandalariga qarshi kurash orqali qishloq xo'jaligi uchun foydali bo'lishi mumkin; ammo, starlings mevalar va unib chiqqan ekinlar bilan oziqlanganida zararkunandalarning o'zi ham bo'lishi mumkin. Oddiy starlings, shuningdek, ularning katta shahar po'stlog'idan kelib chiqqan shovqin va tartibsizlik tufayli bezovtalanishi mumkin. Xususan, kiritilgan populyatsiyalar bir qator nazoratga, shu jumladan yo'q qilish, ammo bular G'arbiy Avstraliyaning mustamlaka qilinishining oldini olishdan tashqari, cheklangan muvaffaqiyatga erishdilar.

1980-yillardan beri shimoliy va g'arbiy Evropaning ayrim qismlarida son kamayib ketdi, chunki o'sayotgan jo'jalar uchun ozuqa sifatida o'tloq umurtqasizlar kam. Shunga qaramay, uning ulkan global aholisi sezilarli darajada kamayib bormoqda deb o'ylamaydilar, shuning uchun oddiy yulduzlar mavjud deb tasniflanadi Eng kam tashvish tomonidan Tabiatni muhofaza qilish xalqaro ittifoqi.

Taksonomiya va sistematikasi

Oddiy starling birinchi marta tasvirlangan Karl Linney uning ichida Systema Naturae hozirgi binomial nomi bilan 1758 yilda.[3] Sturnus va vulgaris dan olingan Lotin navbati bilan "starling" va "common" uchun.[4] Qadimgi ingliz staer, keyinroq qarashva lotin sturnus ikkalasi ham noma'lum narsadan kelib chiqqan Hind-evropa miloddan avvalgi ikkinchi ming yillikka oid ildiz. "Starling" birinchi marta XI asrda qayd etilgan, u erda bu turning balog'at yoshiga etmaganligi haqida so'z yuritilgan, ammo XVI asrga kelib u allaqachon "qarash" ni har qanday yoshdagi qushlarni nazarda tutgan.[5] Eski ismga havola qilinadi Uilyam Butler Yits ' she'riyat "Mening derazam yonidagi yulduzning uyasi".[6] The Xalqaro ornitologik kongress "ingliz tilidagi mahalliy xalq tilidagi ism oddiy starling.[7]

The starling Sturnidae oilasi butunlay Eski dunyo turlarning ko'pligi bilan boshqa joylardan tashqari guruhni Janubi-sharqiy Osiyo va Saxaradan Afrikaga.[8] Jins Sturnus bu polifetetik va uning a'zolari o'rtasidagi munosabatlar to'liq hal qilinmagan. Oddiy yulduzchaning eng yaqin aloqasi bu beg'ubor starling.[9] Ko'chib yurmaydigan beg'ubor yulduz ajdodlari populyatsiyasidan kelib chiqishi mumkin S. vulgaris ichida omon qolgan Iberiya refugium davomida Muzlik davri chekinish,[10] va mitoxondrial gen tadqiqotlar shuni ko'rsatadiki, uni oddiy yulduzchaning pastki turi deb hisoblash mumkin. Oddiy yulduzli populyatsiyalar orasida nomaqbul oddiy yulduzlar va beg'ubor yulduzlar orasida ko'proq genetik o'zgarish mavjud.[11] Oddiy starling qoldiqlari ma'lum bo'lsa-da O'rta pleystotsen,[12] Sturnidae-dagi munosabatlarni hal qilishdagi muammoning bir qismi - bu butun oila uchun tosh qoldiqlarining kamligi.[10]

Subspecies

Bir nechtasi bor pastki turlari o'zgaruvchan oddiy yulduzcha oxir-oqibat kattalar shilliq qavatining rangi va ohangida. Geografik diapazon bo'yicha bosqichma-bosqich o'zgarish va keng qamrovli intergradatsiya shuni anglatadiki, turli xil pastki turlarni qabul qilish hokimiyat o'rtasida farq qiladi.[13][14]

Subspecies[a]
SubspeciesVakolatOraliqIzohlar
Evropalik yulduzcha
(S. v. Vulgaris)
Linney, 1758 yilEvropaning aksariyat qismi, uzoq shimoli-g'arbiy va janubi-sharqdan tashqari; shuningdek Islandiya va Kanareykalar orollariThe subspecies nomzodini ko'rsatish.
Farer boqmoqda
(S. v faroensis)
Feilden, 1872The Farer orollariNomzodlardan biroz kattaroq, ayniqsa qonun loyihasida va oyoqlarda. Qorong'i va xira yashil rangga ega bo'lgan kattalar va hatto yangi tuklarda ham juda kam dog 'bor. Voyaga etmagan odamning sootli oqi, iyagi oqargan va qorindagi joylar; tomoq qoralangan.
Shetland chaqmoqday
(S. v. Zetlandicus)
Xartert, 1918.The Shetland orollariYoqdi faroensis, lekin o'sha kichik turlar orasidagi oraliq va vulgaris. Qushlar Fair Orol, Sent-Kilda va Tashqi gibridlar bilan ushbu subspecies va nominatsiya va joylashtirish o'rtasida oraliqdir vulgaris yoki zetlandikus vakolatiga ko'ra farq qiladi.
Azor yulduzlari
(S. v granti)
Hartert, 1903 yilThe Azor orollariNomzod kabi, lekin kichikroq, ayniqsa oyoqlar. Ko'pincha yuqori qismlarda kuchli binafsha porloq.
Sibir yulduzi
(S. v. Poltaratskiy)
(Finch, 1878)Sharqiy Boshqirdiston sharqqa qarab Ural tog'lari va markaziy Sibir, ga Baykal ko'li va g'arbiy Mo'g'ulistonNomzod kabi, lekin boshida porloq rang asosan pushti rangda, orqa yashil rangda, qanotlarda odatda binafsha-ko'k, yuqori qanot qopqog'ida mavimsi-yashil rangda. Parvoz paytida, qanot pardalari va qo'ltiq osti qismlariga ko'zga tashlanadigan engil darchin-buff qirralari; yangi tuklar ichida bu joylar juda oqarib ko'rinishi mumkin.
Qora dengiz sho'ng'idi
(S. v. Taurikus)
Buturlin, 1904Kimdan Qrim va sharqida Dnepr daryosi sohillari atrofida sharqqa qarab Qora dengiz g'arbga Kichik Osiyo. Uning o'rnini egallagan baland tog'larda topilmadi binafsha rang.Nomzodga o'xshab, ammo qat'iy qanotli. Yashil boshning tanasi, tanasi bronza-binafsha rang, yonboshlari va yuqori qanot qoplamalari yashil-bronza. Yostiqlarning oqargan chekkalari bilan qoraygan. Tuklarni ko'paytirishda deyarli beg'ubor.
Sharqiy Turkiya starling
(S. v. Purpurascens)
Gould, 1868Sharqiy Turkiyaga Tbilisi va Sevan ko'li, Qora dengizning sharqiy qirg'og'idagi tog'larda, o'rnini bosuvchi taurikusNomzodlar singari, lekin qanotlari uzunroq va yashil porloq, quloq pardalari, bo'yin va yuqori ko'krak bilan cheklangan. Binafsharang porlash boshqa joylarda, qanotlarning yuqori qismida, bronza ko'proq. Qopqoqlarga ingichka oq chekkalari bilan qorong'i pastki qavat.
Kavkaz yulduzlari
(S. v. Caucasicus)
Lorenz, 1887The Volga deltasi sharq orqali Kavkaz va qo'shni hududlarBosh va orqa qismida yashil porlash, bo'yin va qorin qismida binafsha rang porlashi, yuqori qanot pardalarida esa mavimsi. Underwings shunga o'xshash binafsha rang.
Markaziy Osiyo starling
(Porfironotus S.)
(Sharpe, 1888)G'arbiy Markaziy Osiyo, ichiga baho berish poltaratskiy o'rtasida Jungari Alatau va Oltoy tog'lariGa juda o'xshash taurikus, lekin kichikroq va to'liq allopatrik, tomonidan ajratilgan binafsha rang, Kavkaz va nobilior.
Xyum boqmoqda
yoki afg'on starling
(S. v. Nobilior)
(Xum, 1879)Afg'oniston, janubi-sharqiy Turkmaniston va qo'shni O'zbekiston sharqqa EronYoqdi binafsha rang, lekin kichikroq va qanotlari qisqaroq; quloq pardalari binafsha rangda, pastki va yuqori qismida esa juda qizg'ish rang bor edi.
Himoloy yulduzlari
(S. v. Humii)
(Bruks, 1876)Kashmir ga NepalKichik; binafsha rang porlashi bo'yin qismida, ba'zida dumaloq plashlarda, aks holda yashil rangda cheklangan. Bu ba'zan nom ostida davolanadi indikus tomonidan berilgan Xojson.[b]
Sindh qaragancha
(Kichik v)
(Xyum, 1873)PokistonKichik; bosh va pastki qorin va orqada cheklangan yashil porloq, aks holda binafsha rangda porloq.

Qushlar Fair Orol, Sent-Kilda va Tashqi gibridlar orasidagi oraliqdir S. v. Zetlandicus va nomzodlar shakli va ularning pastki turlarini joylashtirish vakolatga ko'ra farq qiladi. Ushbu orol shakllariga xos bo'lgan qorong'u balog'atga etmagan bolalar vaqti-vaqti bilan materik Shotlandiyada va boshqa joylarda uchraydi, bu ularning ba'zilarini ko'rsatmoqda gen oqimi dan faroensis yoki zetlandikus, ilgari izolyatsiya qilingan deb hisoblangan pastki turlari.[18][19]

Boshqa bir nechta pastki nomlar berilgan, ammo ular endi haqiqiy emas. Ularning aksariyati turli xil pastki turlarning birlashadigan joylarida yuzaga keladigan darajalardir. Bunga quyidagilar kiradi: S. v. ruthenus Menzbier, 1891 va S. v. Jitkovi Buturlin, 1904, ular orasidagi intergradlar vulgaris va poltaratskiy g'arbiy Rossiyadan; S. v. Graecus Tschusi, 1905 va S. v. Balcanicus Buturlin va Harms, 1909, ular orasidagi intergradlar vulgaris va taurikus janubdan Bolqon markazga Ukraina va butun Yunonistongacha Bosfor; va S. v.Genrichi Stresemann, 1928, o'rtasida intergrad Kavkaz va nobilior shimoliy Eronda. S. v. Persepolis Ticehurst, 1928 yil janubiy Erondan (Fors viloyati ) juda o'xshash vulgarisva bu aniq aholi yoki oddiygina Evropaning janubi-sharqidan kelgan muhojirlarmi, aniq emas.[14]

Tavsif

Voyaga etmagan
Londonda stolga o'tirgan yosh voyaga etmagan bola. Uning tuklari asosan kulrang-jigarrang rangga ega

Oddiy starlingning uzunligi 19-23 sm (7.5-9.1 dyuym), qanotlari 31-44 sm (12-17 dyuym) va vazni 58-101 g (2.0-3.6 oz).[15] Standart o'lchovlar orasida qanotli akkord 11,8 dan 13,8 sm gacha (4,6 dan 5,4 dyuymgacha), quyruq 5.8 dan 6.8 sm gacha (2.3 dan 2.7 dyuymgacha), jinoyatchilar 2,5 dan 3,2 sm gacha (0,98 dan 1,26 gacha) va tarsus 2,7 dan 3,2 sm gacha (1,1 dan 1,3 gacha).[15] The tuklar bu iridescent qora, porloq binafsha yoki yashil rangga bo'yalgan va oq rang bilan to'qilgan, ayniqsa qishda. Voyaga etgan erkak oddiy yulduzchalarning pastki qismlari yilning ma'lum bir davrida kattalar ayollariga qaraganda kamroq aniqlanadi. Erkaklarning tomoq patlari uzun va bo'shashgan bo'lib, ular ko'rgazmada ishlatiladi, urg'ochilarniki esa kichikroq va o'tkirroq. Oyoqlari qattiq va pushti yoki kulrang-qizil rangga ega. Hisob-kitob tor va konus shaklida o'tkir uchi bilan; qishda u jigarrang-qora rangga ega, ammo yozda urg'ochilar limonli sariq tumshuqlarga ega, erkaklar esa ko'k-kulrang poydevorli sariq qog'ozga ega. Moulting yiliga bir marta - naslchilik mavsumi tugagandan so'ng yoz oxirida sodir bo'ladi; yangi tuklar ko'zga ko'rinadigan oq (ko'krak patlari) yoki buff (qanot va orqa tuklar) bilan uchirilgan bo'lib, bu qushga dog 'ko'rinishini beradi. Naslchilik davridagi dog 'miqdorining pasayishiga oq tuklar uchlari asosan eskirgan holda erishiladi. Voyaga etmaganlar kulrang-jigarrang va birinchi qishdayoq kattalarga o'xshaydilar, lekin ko'pincha jigarrang tuklarni saqlab qolishmoqda, ayniqsa boshlarida.[13][20] Ular odatda irislarning rangi bilan, erkaklarda boy jigarrang, ayollarda sichqon-jigarrang yoki kul rang bilan jinsiy aloqa qilishlari mumkin. An o'rtasidagi ziddiyatni taxmin qilish ìrísí va markaz har doim qorong'i o'quvchi jinsiy aloqani aniqlashda 97% aniq, agar tomoq patlarining uzunligi ham hisobga olinsa, 98% gacha ko'tariladi.[21][22] Oddiy starling ham starling standartlari, ham passerin standartlari bo'yicha o'rtacha kattalikka ega. Kabi boshqa o'rta kattalikdagi passerinlardan osongina ajralib turadi itlar, icteridlar yoki kichik koridlar, nisbatan qisqa dumidan, o'tkir, pichoqqa o'xshash velosipeddan, dumaloq qorin shaklidan va kuchli, katta (va rangdor) oyoqlardan. Parvoz paytida uning kuchli uchli qanotlari va qorong'i ranglari ajralib turadi, yerda uning g'alati, biroz bemalol yurishi ham xarakterlidir. Bo'yash va qurish odatda bu qushni boshqa yulduzchalardan ajratib turadi, garchi ular bir-biri bilan chambarchas bog'liq bo'lsa ham beg'ubor starling jismonan kattalar naslchilik shilliqlarida iridescent dog'lar etishmasligi bilan ajralib turishi mumkin.[23]

Voyaga etmaganlar uchun moulting
Kaliforniyada voyaga etmagan. U qisman o'zining birinchi qishki shilliq qavatiga qo'shilgan; shu bilan birga, balog'at yoshiga etmagan jigarrang tuklar bosh va bo'yin qismida taniqli

Ko'pgina quruqlik singari oddiy yulduzlar sakrashdan ko'ra yurish yoki yugurish bilan harakat qilishadi. Ularning parvozi juda kuchli va to'g'ridan-to'g'ri; ularning uchburchak shaklidagi qanotlari juda tez urishadi va vaqti-vaqti bilan qushlar qisqa masofani bosib o'tib, baland parvozni davom ettirishdan oldin balandlikni yo'qotmaydi. Qushlar suruvda deyarli bir vaqtning o'zida uchishadi, g'ildirakda aylanadilar va bir ovozdan aylanadilar, ixcham massani hosil qiladilar yoki aqlli oqimga o'tadilar, yana to'planib, kelishilgan holda erga tushadilar.[20] Ko'chib yurishdagi oddiy yulduzcha soatiga 60-80 km (37-50 milya) tezlikda uchib, 1000–1500 km (620–930 milya) masofani bosib o'tishi mumkin.[24]

Bir nechta quruqlikdagi yulduzlar, shu jumladan tur Sturnus, proba bilan oziqlantirishga yordam beradigan bosh suyagi va mushaklarning moslashuvi mavjud.[25] Ushbu moslashuv oddiy yulduzchada eng kuchli rivojlangan (bilan birga beg'ubor va oq yonoqli starlings ), qaerda protraktor mushaklari jag'ning ochilishi uchun mas'ul kattalashtirilgan va bosh suyagi tor bo'lib, ko'zni oldinga siljitish uchun imkon beradi.[26] Ushbu texnikaga hisobni erga kiritish va uni yashirin oziq-ovqat mahsulotlarini qidirish usuli sifatida ochish kiradi. Oddiy starlings bu ovqatlanish usulidan foydalanishga imkon beradigan jismoniy xususiyatlarga ega, bu shubhasiz turlarning uzoqqa tarqalishiga yordam berdi.[15]

Iberiyada, g'arbiy O'rta er dengizi va Afrikaning shimoli-g'arbiy qismida oddiy yulduzchani bir-biri bilan chambarchas bog'liq bo'lgan beg'ubor yulduzcha bilan aralashtirib yuborish mumkin, uning tuklari, nomi bilan aytganda, bir xilroq rangga ega. Yaqin masofadan ko'rinib turibdiki, ikkinchisining tomoq tuklari uzunroq, bu haqiqatan ham u qo'shiq aytganda seziladi.[27]

Vokalizatsiya

Guruhning suhbatlari
Voyaga etgan erkak
Voyaga etgan erkak qo'shiq aytmoqda va uzun tomoq patlarini namoyish qilmoqda
Qo'shiq Sturnus vulgaris, Spring Creek Park, Nyu-York, AQSh.

Oddiy starling shovqinli qushdir. Uning Qo'shiq tovushlarning marosim ketma-ketligi tarkibida melodik va mexanik-tovushli shovqinlarning xilma-xilligidan iborat. Erkak asosiy qo'shiqchi bo'lib, bir daqiqa yoki undan ko'proq davom etadigan qo'shiqlar bilan shug'ullanadi. Ularning har biri odatda to'rt xil qo'shiq turini o'z ichiga oladi, ular bir-birlarini pauzasiz muntazam ravishda kuzatib boradilar. Bahs sof ohangdor hushtak chalishlaridan boshlanadi va shundan keyin qo'shiqning asosiy qismi, ko'pincha qo'shiqning ovozini o'z ichiga olgan bir qator o'zgaruvchan ketma-ketliklar qo'shiladi. taqlid qildi qushlarning boshqa turlaridan va turli xil tabiiy yoki sun'iy shovqinlardan. Taqlid qilingan tovushning tuzilishi va soddaligi uning paydo bo'lish chastotasidan ko'ra muhimroqdir. Ba'zi hollarda yovvoyi yulduzcha bir marta eshitgan tovushni taqlid qilishi kuzatilgan. Har bir tovush klipi qush ikkinchisiga o'tishdan oldin bir necha marta takrorlanadi. Ushbu o'zgaruvchan qismdan keyin bir necha marta bosilgan bir necha marta bosish va keyin bir necha turdagi shakllangan yuqori chastotali qo'shiqlarning so'nggi portlashi keladi. Har bir qushning o'ziga xos xususiyati bor repertuar 35 ga qadar o'zgaruvchan qo'shiq turlari va 14 marta bosish imkoniyatiga ega bo'lgan yanada malakali qushlar bilan.[28]

Ko'payish davri yaqinlashganda erkaklar doimiy ravishda qo'shiq aytadilar va juftlik bog'langandan keyin kamroq bajaradilar. Urg'ochi oldida ba'zida erkak o'z uyasiga uchib kirib, eshikdan qo'shiq aytadi, aftidan ayolni o'ziga jalb qilishga urinadi. Keksa qushlar yoshlariga qaraganda kengroq repertuarga ega. Uzunroq qo'shiq kuylash bilan shug'ullanadigan va repertuarlari kengroq bo'lgan erkaklar turmush o'rtoqlarni ilgari jalb qilishadi va ko'proq reproduktiv muvaffaqiyat boshqalarga qaraganda. Ayollar murakkabroq qo'shiqlari bilan turmush o'rtog'ini afzal ko'rishadi, ehtimol bu katta tajriba yoki uzoq umrni ko'rsatishi mumkin. Murakkab qo'shiqqa ega bo'lish, shuningdek, hududni himoya qilish va kam tajribali erkaklarni tajovuzdan saqlashda foydalidir.[28]

Bosh suyagi va mushaklarning qo'shiq aytish uchun moslashuvi bilan bir qatorda erkak yulduzchalari ham kattaroqdir sirinx ayollarga qaraganda. Bu mushak massasining ko'payishi va shprits skeletining kattalashgan elementlari bilan bog'liq. Erkak yulduzchasi siringi ayol hamkasbidan 35 foizga katta.[29] Ammo, bu jinsiy dimorfizm, zebra finch singari qo'shiq qushlari turlariga qaraganda kamroq seziladi, bu erda erkaklar sirini ayolning sirinkasidan 100% kattaroqdir.[30]

Qo'shiq, naslchilik davridan tashqari, butun yil davomida sodir bo'lgan naslchilik davridan tashqarida ham sodir bo'ladi. Qo'shiqchilar ko'proq erkaklar, ammo ayollar ham vaqti-vaqti bilan qo'shiq aytishadi. Bunday mavsumdan tashqari qo'shiqning vazifasi juda yaxshi o'rganilmagan.[28] Qo'ng'iroqning yana 11 turi tasvirlangan, ular orasida suruv, tahdid, hujum, qo'ng'iroq va kopulyatsiya chaqiruvlari mavjud.[31] Budilnik qo'ng'irog'i qattiq qichqiriqdir va oddiy yulduzlar birlashganda tinimsiz janjal qilishadi.[20] Ular xo'roz paytida va cho'milish paytida suhbatlashishadi, bu juda ko'p shovqinni keltirib chiqaradi, bu esa yaqin atrofda yashovchi odamlarning tirnash xususiyati keltirib chiqarishi mumkin. Oddiy starlinglar to'plami birgalikda uchayotganda, qushlarning qanotlari sinxronlashtirilgan harakatlari yuzlab metr masofada eshitiladigan o'ziga xos ovozni chiqaradi.[31]

Xulq-atvor va ekologiya

Katta bir suruv Rotterdam, Niderlandiya

Oddiy starling, ayniqsa kuz va qishda juda g'aroyib tur. Garchi suruv hajmi juda o'zgaruvchan, ulkan, shovqinli suruvlar - shovqinlar - uylar yaqinida paydo bo'lishi mumkin. Ushbu zich qushlarning konsentratsiyasi hujumlardan himoya deb o'ylashadi yirtqich qushlar kabi peregrine lochinlari yoki Evroosiyo chumchuqlari.[32][33] Floclar mahkamlashadi soha - uchish paytida shakllanish kabi, tez-tez kengayib boruvchi va qisqaradigan va o'zgaruvchan shakl, ko'rinishda har qanday etakchisiz. Har bir oddiy yulduzcha eng yaqin qo'shnilarining harakati natijasida o'z yo'nalishini va tezligini o'zgartiradi.[34]

1,5 milliongacha bo'lgan juda katta xo'rozlar shahar markazlarida, o'rmonzorlarda va qamishzorlarda hosil bo'lib, ularning axlati bilan bog'liq muammolarni keltirib chiqaradi. Ular 30 sm (12 dyuym) gacha chuqurlikda to'planib, kimyoviy moddalarning konsentratsiyasi bilan daraxtlarni o'ldirishi mumkin. Kichikroq miqdordagi axlat a vazifasini bajaradi o'g'it va shuning uchun o'rmon xo'jaligi menejerlari tuproqni yaxshilashdan foyda olish va katta zaharli qatlamlardan saqlanish uchun daraxtlarni bir maydondan boshqasiga ko'chirishga harakat qilishlari mumkin.[35]

Bir milliondan ortiq oddiy yulduzlarning suruvlari janubi-g'arbiy qismida bahorda quyosh botishidan oldin kuzatilishi mumkin Yutland, Daniya dengiz qirg'og'ida botqoq erlar ning Tonder va Esbjerg Tønder va orasidagi munitsipalitetlar Ribe. Ular mart oyida shimoliy Skandinaviya qushlari aprel oyining o'rtalariga qadar ko'payish uchun ketguncha yig'ilishadi. Ularning to'daning harakati osmonga siluetli murakkab shakllarni yaratadi, bu hodisa mahalliy sifatida tanilgan saralash ("qora quyosh").[36]Beshdan ellik minggacha oddiy yulduzchadan iborat har qanday narsaning suruvlari Buyuk Britaniyaning qish o'rtalarida quyosh botishidan oldin paydo bo'ladi. Ushbu suruvlar odatda mızıltılar deb nomlanadi.[37]

Oziqlantirish

Shimoliy Irlandiyadagi fermada boqilayotgan suruv

Oddiy starling asosan hasharotlarga qarshi va zararkunandalarga ham, boshqalarga ham oziklanadi artropodlar. Oziq-ovqat turlari qatoriga kiradi o'rgimchaklar, kran chivinlari, kuya, chivinlar, ninachilar, qizaloq uchadi, chigirtkalar, quloqchinlar, iplar, caddisflies, chivinlar, qo'ng'izlar, arra chivinlari, asalarilar, ari va chumolilar. Yirtqichlar kattalar va lichinkalar rivojlanish bosqichida iste'mol qilinadi va oddiy yulduzchalar ham ovqatlanadilar yomg'ir qurtlari, shilliq qurtlar, kichik amfibiyalar va kaltakesaklar.[38] Iste'mol qilish paytida umurtqasizlar muvaffaqiyatli naslchilik uchun zarur, oddiy starlings hamma narsaga yaroqli va shuningdek eyishi mumkin donalar, urug'lar, mevalar, nektar va oziq-ovqat chiqindilari agar imkoniyat bo'lsa.[39][40][41] Sturnidae ko'pgina qushlardan ajralib turadi, chunki ular tarkibida ko'p miqdordagi ovqatlarni osonlikcha metabolizm qila olmaydi saxaroza, garchi ular uzum va gilos kabi boshqa mevalarni engishsa ham.[42] Oddiy starlingning ajratilgan Azores kichik turlari yo'qolib ketish xavfi ostida bo'lganlarning tuxumlarini iste'mol qiladi roseate tern. Bahorda naslchilik koloniyalariga qaytib kelguniga qadar qirg'in qilish orqali oddiy starling populyatsiyasini kamaytirish choralari ko'rilmoqda.[11]

Voyaga etgan yosh jo'jalarni boqish va ovqat topish

Oddiy chakalakzorlar oziq-ovqat mahsulotlarini olishning bir qancha usullari mavjud, ammo aksariyat hollarda ular hasharotlarni er ostidan yoki ostidan olib, erga yaqin ozuqa olishadi. Odatda, oddiy yulduzlar qisqa o'ralgan o'tlar orasida yem-xashak qilishni afzal ko'rishadi va yaylov hayvonlar bilan yoki orqa tomonidagi perch bilan ovqatlanishadi,[41] bu erda ular sutemizuvchilarning tashqi parazitlari bilan oziqlanadi.[15] Katta podalar "rollarda boqish" deb nomlanuvchi amaliyotni boshlashi mumkin, bu erda suruvning orqa qismidagi qushlar doimiy ravishda ovqatlanish imkoniyatlari eng yaxshi tomonga uchib ketishadi.[39] Suruv qanchalik katta bo'lsa, ozuqa paytida bir-biriga yaqinroq bo'lishadi. Flocks ko'pincha bir joyda bir muncha vaqt ovqatlanadilar va oldingi muvaffaqiyatli oziqlangan saytlarga qaytadilar.[39]

Oddiy starlingda uch xil ovqatlanish harakati kuzatiladi. "Zondlash" qushni hasharot topilmaguncha tasodifiy va takroriy ravishda erga singdirishni o'z ichiga oladi va ko'pincha qush tuynukni kattalashtirish uchun tuproqdagi tumshug'ini ochadigan joyda bo'sh joy bilan birga keladi. Dastlab tasvirlangan ushbu xatti-harakatlar Konrad Lorenz va nemischa atamani hisobga olgan holda zirkeln,[43] plastik axlat qoplarida teshiklarni yaratish va kengaytirish uchun ham ishlatiladi. Ushbu usulni takomillashtirish uchun yosh oddiy chakalakzorlar uchun vaqt talab etiladi va shu sababli yosh qushlarning parhezida ko'pincha kamroq hasharotlar bo'ladi.[27] "Xoking "bu uchayotgan hasharotlarni to'g'ridan-to'g'ri havodan olish," o'pish "- bu harakatni ushlab turish uchun oldinga urishning kamroq tarqalgan usuli. umurtqasizlar yerda. Yomg'ir qurtlari tuproqdan tortib olinmoqda.[39] Ovqatga kirish imkoni bo'lmagan yoki oziqlanish uchun yorug'lik vaqtini qisqartirgan oddiy starlings tanadagi massani yog 'tushishi bilan qoplaydi.[44]

Uyalash

Angliyada daraxt teshigidagi uyasida jo'jasini boqayotgan ota-ona

Juftlik qilmagan erkaklar mos bo'shliqni topib, yolg'iz ayollarni jalb qilish uchun uyalar qurishni boshlaydilar, ko'pincha uyani gullar va yangi yashil materiallar kabi bezaklar bilan bezashadi, keyinroq ayol uni turmush o'rtog'i sifatida qabul qilgandan keyin uni qismlarga ajratadi.[31][45] Yashil materialning miqdori muhim emas, chunki ba'zilari mavjud bo'lsa-da, ammo mavjud giyohlar dekorativ materialda turmush o'rtog'ini jalb qilishda muhim ahamiyatga ega ko'rinadi. Kabi o'simliklarning hidlari yarrow vazifasini bajaradi hid ayollarga jalb qiluvchi.[45][46]

Erkaklar qurilishning ko'p qismida va hatto ayol o'z uyasiga yaqinlashganda ham qo'shiq aytishadi. Keyingi ko'paytirish, erkak va ayol uyani qurishda davom etmoqda. Uyalar har qanday teshikda bo'lishi mumkin, keng tarqalgan joylarga ichi bo'sh daraxtlar, binolar, daraxtlar stuntlari va sun'iy uyalar kiradi.[31] S. v. Zetlandicus odatda jarliklarda va jarlikdagi teshiklarda ko'payadi, yashash joyi faqat nomzod shaklida kamdan kam qo'llaniladi.[47] Uyalar odatda patlar, jun va yumshoq barglardan tashkil topgan ichki qoplamali somon, quruq o't va novdalardan tayyorlanadi. Qurilish odatda to'rt yoki besh kun davom etadi va inkubatsiya orqali davom etishi mumkin.[31]

Oddiy starlings ikkalasi ham monogam va ko'pxotinli; garchi zotlar odatda bitta erkak va bitta ayol tomonidan tarbiyalangan bo'lsa-da, ba'zida juftlikda qo'shimcha yordamchi bo'lishi mumkin. Juftliklar koloniyaning bir qismi bo'lishi mumkin, bu holda bir nechta boshqa uyalar bir xil yoki yaqin atrofdagi daraxtlarni egallashi mumkin.[31] Erkaklar ikkinchi urg'ochi bilan juftlashishi mumkin, birinchisi hali uyasida. Qushlarning reproduktiv muvaffaqiyati ikkinchi uyada birlamchi uyaga qaraganda yomonroq va erkak monogam bo'lib qolganda yaxshi bo'ladi.[48]

Naslchilik

Uyadagi beshta tuxum
Tuxum, to'plam Visbaden muzeyi, Germaniya
Oziqlanishni kutayotgan jo'jalar
Uyasining kirish qismida ovqatlanishni kutayotgan jo'jalar devor oralig'ida joylashgan Geyvey, Irlandiya

Naslchilik bahor va yoz oylarida amalga oshiriladi. Kopulyatsiyadan so'ng, ayol bir necha kun davomida har kuni tuxum qo'yadi. Agar bu vaqt ichida tuxum yo'qolsa, u uni almashtirish uchun boshqasini qo'yadi. Odatda to'rt yoki beshta tuxum mavjud ovoid shakli va och ko'k yoki vaqti-vaqti bilan oq rangga ega bo'lib, ular odatda porloq ko'rinishga ega.[31] Tuxumlarning rangi past nur darajasida ko'k rangning nisbatan yaxshi ko'rinishi orqali rivojlanganga o'xshaydi.[49] Tuxum kattaligi uzunligi 26,5-34,5 mm (1,04-1,36 dyuym) va maksimal diametri 20,0-22,5 mm (0,79-0,89 dyuym).[15]

Kuluçka o'n uch kun davom etadi, garchi oxirgi qo'yilgan tuxum birinchi chiqqanidan 24 soat ko'proq vaqt talab qilishi mumkin. Ikkala ota-ona ham tuxumni parvarish qilish mas'uliyatini o'z zimmalariga olishadi, ammo urg'ochi ularni erkaklarnikiga qaraganda ko'proq inkubatsiya qilish uchun sarflaydi va tunda erkak kommunal xo'rozga qaytib kelganda buni qiladigan yagona ota-ona. Yoshlar ko'r va yalang'och tug'ilishadi. Ulardan chiqqandan keyin etti kun ichida ular mayin paxmoq bo'lib rivojlanib, to'qqiz kun ichida ko'rishlari mumkin.[31] Jo'jalar tana haroratini tartibga sola olgandan so'ng, olti kundan keyin,[50] kattalar uyadan axlatni olib tashlashni deyarli to'xtatadilar. Bundan oldin, ifloslanish jo'jalarning tuklarini ham, uyadagi materiallarni ham namlaydi va shu bilan ularning izolyatsiyasi sifatini pasaytiradi va lyuklarni sovutish xavfini oshiradi.[51] Nestlings uch hafta davomida uyada qoling, u erda ikkala ota-ona tomonidan doimiy ravishda boqiladi. Fleglings yana bir yoki ikki hafta davomida ota-onalari tomonidan ovqatlanishni davom eting. Bir juftlik yiliga uchta bolani boqishi mumkin, ko'pincha bitta uyani qayta ishlatadi va qaytadan bog'lanadi,[31] ikkita zoti odatiy bo'lsa ham,[15] yoki 48 ° shimoldan faqat bir shimolda.[24] Ikki oy ichida ko'pchilik balog'at yoshiga etmaganlar va birinchi tuklarni olishadi. Ular keyingi yil kattalar shilliq qavatini sotib olishadi.[31] Boshqa passerinlarda bo'lgani kabi, uya toza va jo'jalar ' najas torbalari kattalar tomonidan olib tashlanadi.[52]

Ichki parazitlar umumiy starling uyalarida keng tarqalgan. Koloniyalarda mavjud bo'lgan urg'ochi "suzuvchi" lar (nasl berish davrida juftlashtirilmagan urg'ochilar) ko'pincha boshqa juft uyasida tuxum qo'yadilar.[53] Bola bolalari o'z uylariga yoki qo'shni uyalariga bostirib kirib, yangi naslni haydab chiqarishi haqida ham xabar berilgan.[31] Oddiy starling uyalari 48% dan 79% gacha muvaffaqiyatli uchish darajasiga ega, garchi uloqchalarning atigi 20% i nasl yoshiga qadar omon qoladi; kattalar hayot darajasi 60% ga yaqin. O'rtacha umr ko'rish davomiyligi 2-3 yil,[24] 22 yr 11 m uzoq umr ko'rish rekordi bilan.[54]

Yirtqichlar va parazitlar

Yulduzli yirtqichlarning aksariyati qushlardir. Starling guruhlarining odatiy munosabati parvoz qilishdir, odatdagidek tez va tezkor naqshlarda baland uchadigan yulduz yulduzlarining to'lqinlari. Ularning parvozdagi qobiliyatlari kamdan-kam yirtqich qushlarga to'g'ri keladi.[55][56] Voyaga etgan oddiy starlings ovlanadi qirg'iylar kabi shimoliy goshawk (Accipiter gentilis) va Evroosiyo chumchuqlari (Accipiter nisus),[57] va lochinlar shu jumladan peregrine lochin (Falco peregrinus), Evroosiyo xobbi (Falco subbuteo) va oddiy kestrel (Falco tinnunculus).[58][59] Sekinroq raptors yoqadi qora va qizil kites (Milvus migrans va milvus), sharqiy imperiya burguti (Aquila heliaca), oddiy shov-shuv (Buteo buteo) va Australasian harrier (Sirk taxminiylari) osonroq ushlanadigan qushchalarni yoki balog'atga etmagan bolalarni olishga moyil.[60][61][62] Kechasi guruhlarda o'tirganlarida, ular boyqushlarga, shu jumladan kichik boyqush (Afina noctua), uzun quloqli boyqush (Asio otus), kalta quloqli boyqush (Asio flammeus), boyqush (Tyto alba), jo'xori boyo'g'li (Strix aluco) va Evroosiyo burgut-boyo'g'li (Bubo bubo).[63][64]

Yigirmadan ortiq turlari qirg'iy, boyqush va lochin Shimoliy Amerikada vaqti-vaqti bilan yirtqich yulduzchalardan oldin paydo bo'lganligi ma'lum, ammo kattalar orasida eng odatiy yirtqichlar shaharliklar bo'lishi mumkin. peregrine lochinlari yoki merlinlar (Falco columbarius).[65][66] Oddiy mynalar (Acridotheres tristis) ba'zida tuxum, uyalay va katta yoshli yulduzchalarni uyalaridan haydab chiqaradi,[31] va kamroq asal uchun qo'llanma (Kichik ko'rsatkich), a parazit, oddiy starlingni xost sifatida ishlatadi.[67] Starlings ko'proq aybdorlar, lekin uyni haydash qurbonlari emas, lekin, ayniqsa, boshqa starlings va qarag'aylar.[68][69] Ularga ko'tarilishga qodir sutemizuvchilar, masalan, uyalarga bostirib kirishi mumkin stullar (Mustela erminea), rakunlar (Procyon lotor)[70][71] va sincaplar (Sciurus spp.),[24] va mushuklar beparvolarni tutishi mumkin.[72]

Oddiy starlings - parazitlarning keng doirasi. AQShning oltita shtatidan uch yuzta oddiy yulduzcha ishtirokida o'tkazilgan so'rov natijalariga ko'ra, ularning barchasida parazitning kamida bitta turi borligi aniqlandi; 99 foizida tashqi burgalar, oqadilar yoki shomillar bo'lgan, 95 foizida esa ichki parazitlar, asosan qurtlarning har xil turlari bo'lgan. Qon so'ruvchi turlar o'lganida o'z uy egasini tark etadi, ammo boshqa tashqi parazitlar murda ustida qoladi. Hisob shakli deformatsiyalangan qushga juda ko'p zarar etkazildi Mallofaga bitlar, ehtimol uning zararli moddalarni olib tashlay olmasligi sababli.[73]

Parazit oqadilar
Dermanyssus gallinae, oddiy starling paraziti

Tovuq burga (Ceratophyllus gallinae ) eng keng tarqalgan burga ularning uyalarida.[74] Kichkina, rangpar uy-chumchuq burga C. fringillae, shuningdek, vaqti-vaqti bilan u erda uchraydi va, ehtimol, uning asosiy egasi boshqa turlarning uyalarini egallab olish odatidan kelib chiqadi. Ushbu burga AQShda, hatto bundan keyin ham bo'lmaydi uy chumchuqlari.[75] Bitlar o'z ichiga oladi Menacanthus eurystemus, Brueeliya tumanligi va Stumidoecus sturni. Boshqa artropod parazitlari kiradi Ixodlar Shomil va oqadilar kabi Analgopsis passerinus, Boydaia stumi, Dermanyssus gallinae, Ornithonyssus bursa, O. sylviarum, Proktofillalar turlari, Pteronyssoides truncatus va Trouessartia rosteri.[76] Tovuq oqimi D. gallinae o'zi yirtqich oqadilar tomonidan o'lja qilingan Androlaelaps casalis. Parazit turlari sonida ushbu nazoratning mavjudligi qushlarning eski uyalarni qayta ishlatishga tayyorligini tushuntirishi mumkin.[77]

Oddiy yulduzchalarni parazit qiladigan uchuvchi hasharotlarga quyidagilar kiradi chivin Omithomya nigricornis[76] va saprofag pashsha Camus hemapterus. Oxirgi tur egasining patlarini sindirib, tuklar o'sishi natijasida hosil bo'lgan yog'lar bilan yashaydi.[78] Kuya lichinkalari Hofmannophila pseudospretella kabi hayvonot materiallari bilan oziqlanadigan uyni tozalash vositalaridir najas yoki o'lik chaqaloqlar.[79] Protozoy jinsning qon parazitlari Hemoproteus oddiy yulduzchalarda topilgan,[80] ammo taniqli zararkunanda - bu porloq qizil rang nematod Sinxamus traxeyasi. Ushbu qurt o'pkadan to ga o'tadi traxeya va uning mezbonini bo'g'ib qo'yishi mumkin. Britaniyada qal'a va oddiy starling eng ko'p zararlangan yovvoyi qushlardir.[81] Boshqa qayd qilingan ichki parazitlarga tikanli qurt kiradi Protorhynchus transverslari.[82]

Oddiy starlings shartnoma tuzishi mumkin parrandachilik,[83][84] parranda bezgagi[85][86] va retrovirus - tushuntirilgan limfomalar.[87] Asirga olingan starlings ko'pincha jigarda ortiqcha temirni to'playdi, bu esa ovqatga qora choy barglarini qo'shish orqali oldini olish mumkin.[88][89]

Tarqatish va yashash muhiti

Oddiy yulduzchalarning dunyo aholisi 2004 yilda 3,8 million kishidan iborat bo'lib, ularning umumiy maydoni 8,870,000 km ni tashkil etgan.2 (3,420,000 sqm mil).[90] Shimoliy yarim sharda keng tarqalgan, bu qush mahalliy hisoblanadi Evroosiyo va Evropada, shimoliy Afrikada (dan.) joylashgan Marokash ga Misr ), Hindiston (asosan shimolda, lekin muntazam ravishda janubga qarab kengayib boradi)[91] va Maldiv orollariga tarqaldi[92]) Nepal, Yaqin Sharq, shu jumladan Isroil, Suriya, Eron va Iroq va shimoliy-g'arbiy Xitoy.[90]

Migratsiya bo'yicha dam olish
Ko'chish paytida qarag'ay daraxtiga suyanadigan suruv

Evropaning janubi va g'arbiy qismida va 40 ° shimoliy kenglik janubidagi oddiy yulduzlar asosan rezident,[24] boshqa aholi qish qattiq bo'lgan hududlardan ko'chib kelganiga qaramay, er muzlagan va oziq-ovqat tanqisligi. Shimoliy Evropa, Rossiya va Ukrainadan ko'p sonli qushlar janubi g'arbga yoki janubi sharqqa ko'chib yurishadi.[20][28] Kuzda, Sharqiy Evropadan muhojirlar kelganda, Britaniyaning oddiy yulduzlari ko'pchilik yo'lga chiqmoqdalar Iberiya va Shimoliy Afrika. Qushlarning boshqa guruhlari mamlakat bo'ylab o'tmoqda va bu qushlarning turli oqimlari yo'llari kesib o'tishi mumkin.[20] 15000 qushlardan ini kabi qo'ng'iroq qilishdi Mersisayd, Angliya, jismoniy shaxslar yilning turli vaqtlarida Norvegiya, Shvetsiya, Finlyandiya, Rossiya, Ukraina, Polsha, Germaniya va Kam mamlakatlar.[93] Yaponiyada va Gongkongda oddiy yulduzlar kamdan-kam uchraydi, ammo bu qushlar qaerdan paydo bo'lganligi noma'lum.[28] Shimoliy Amerikada shimoliy populyatsiyalar qishda Kanadaning katta qismini bo'shatib, migratsiya tartibini ishlab chiqdilar.[94] Mamlakat sharqidagi qushlar janubga, g'arbiy tomondan esa AQShning janubi-g'arbiy qismida qishlashadi.[15]

Oddiy starlings sun'iy inshootlar va daraxtlar etarli darajada uyalashni ta'minlaydigan shahar yoki shahar atroflarini afzal ko'radi pishirish saytlar. Qamishzorlar ham xo'roz uchun qulaydir va qushlar odatda qishloq xo'jaligi erlari, yaylov yaylovlari, o'yin maydonchalari, golf maydonchalari va aerodromlar kabi o'tloqli joylarda oziqlanadi.[39] Ular vaqti-vaqti bilan ochiq o'rmonlar va o'rmonzorlarda yashaydilar va ba'zida avstraliyaliklar kabi butazor joylarda uchraydi sog'liqni saqlash. Oddiy starlings kamdan-kam hollarda zich, nam o'rmonlarda yashaydi (ya'ni yomg'ir o'rmonlari yoki nam) sklerofil o'rmonlar), ammo qirg'oqbo'yi hududlarida joylashgan bo'lib, u erda uloqlar uyalar va jarliklarda yashaydilar dengiz o'tlari. Ularning turli xil yashash joylariga moslashish qobiliyati ularga tarqalish va dunyoning turli joylarida o'rnashishga imkon berdi, natijada qirg'oqdagi botqoqlardan tortib to yashash joylariga qadar. alp dengiz jarliklaridan to balandlikgacha bo'lgan dengiz sathidan 1900 m (6200 fut) balandlikdagi o'rmonlar.[39]

Kiritilgan populyatsiyalar

Oddiy yulduzcha Yangi Zelandiya, Avstraliya, Janubiy Afrika, Shimoliy Amerika, Fidji va bir necha Karib orollari. Natijada, u ham ko'chib o'tishga muvaffaq bo'ldi Tailand, Janubi-Sharqiy Osiyo va Yangi Gvineya.[39]

Janubiy Amerika

Besh kishi Angliyadan kemaga etkazilgan Lago-de-Marakaybo 1949 yil noyabrda Venesuelada, ammo keyinchalik g'oyib bo'ldi.[95] 1987 yilda shahardagi bog'larda oddiy chakalakzorlarning oz sonli populyatsiyasi kuzatilgan Buenos-Ayres.[41][96] O'shandan beri, yo'q qilish bo'yicha ba'zi bir dastlabki urinishlarga qaramay, qush Atlantika sohilidan 30 km (19 milya) masofada turib, yiliga o'rtacha 7,5 km (4,7 milya) tezlikda ko'payib bormoqda. Argentinada bu tur turli xil tabiiy va sun'iy uyalar joylaridan, xususan, daraxtzor teshiklaridan foydalanadi.[96]

Avstraliya

Oddiy starling Avstraliyaga qishloq xo'jaligi ekinlarining hasharot zararkunandalarini iste'mol qilish uchun kiritilgan. Dastlabki ko'chib kelganlar, ularning paydo bo'lishini intizorlik bilan kutishar, oddiy changalzorlar ham changlanish uchun muhim ahamiyatga ega deb hisoblashgan zig'ir, asosiy qishloq xo'jaligi mahsuloti. Yangi chiqarilgan qushlar uchun uy-qutilar fermer xo'jaliklarida va ekin maydonlariga joylashtirildi. Oddiy starling tanishtirildi Melburn 1857 yilda va Sidney yigirma yil o'tgach.[39] 1880-yillarga kelib, mamlakatning janubi-sharqida iqlimlashtirish qo'mitalarining faoliyati tufayli tashkil etilgan aholi mavjud edi.[97] 20-asrning 20-yillariga kelib, oddiy yulduzchalar keng tarqaldi Viktoriya, Kvinslend va Yangi Janubiy Uels, ammo o'sha paytgacha ular zararkunandalar deb hisoblanardi.[39] Garchi oddiy starlings birinchi marta ko'rilgan bo'lsa Albani, G'arbiy Avstraliya 1917 yilda ular asosan davlatga tarqalishining oldi olindi. Keng va quruq Nullarbor tekisligi tabiiy to'siqni ta'minlaydi va o'ttiz yil ichida 55000 qushni o'ldirgan kurash choralari ko'rildi.[98] Oddiy starling ham mustamlakaga aylandi Kenguru oroli, Lord Xou oroli, Norfolk oroli va Tasmaniya.[95]

Yangi Zelandiya

Yangi Zelandiyadagi dastlabki ko'chmanchilar tupni tozalashdi va yangi ekilgan ekinlarni tırtıllar va boshqa hasharotlar avvalgi oziq-ovqat manbalaridan mahrum bo'lganlar. Mahalliy qushlar odamga yaqin joyda yashashga odatlanmaganlar, shuning uchun oddiy starling Evropadan ham keltirilgan Uy chumchuq zararkunandalarga qarshi kurashish. U birinchi bo'lib 1862 yilda Nelson iqlimlashtirish jamiyati tomonidan olib kelingan va boshqa tanishtiruvlar amalga oshirilgan. Tez orada qushlar tashkil topdi va endi butun mamlakat bo'ylab, shu jumladan subtropikada ham uchraydi Kermadek orollari shimolga va bir xil masofada joylashgan Makquari oroli uzoqroq janubda.[99][100]

Shimoliy Amerika

Yulduzli baliqlar
Flock in Napa vodiysi, Kaliforniya
Da Yarim oy ko'rfazi, Kaliforniya
Missuri shtatining Sent-Luis shahri atrofida parvoz qilayotgan yevropalik Starling.

Ikki muvaffaqiyatsiz urinishdan so'ng,[101] 1890 yilda Nyu-Yorkka 60 ga yaqin oddiy starlings chiqarildi Markaziy Park tomonidan Evgeniy Sxeffelin. U prezident edi Amerika Iqlimlashtirish Jamiyati xabarlariga ko'ra, bu asarlarda aytib o'tilgan har bir qush turini tanishtirishga harakat qilgan Uilyam Shekspir Shimoliy Amerikaga,[102][103] garchi bu bahsli bo'lsa ham.[104][105] Xuddi shu sanada Portlend Song Bird Club Oregon shtatining Portlend shahrida 35 juft oddiy starlingsni ozod qildi. Ushbu qushlar tashkil topgan, ammo 1902 yil atrofida g'oyib bo'lgan. 1940-yillarning o'rtalarida Tinch okeanining shimoli-g'arbiy qismida oddiy yulduzlar paydo bo'lgan va bu qushlar, ehtimol, 1890 yilgi Markaziy Park kirishining avlodlari bo'lgan.[101] Dastlab 60 ta qush 150 millionga ko'payib, Kanadaning janubi va Alyaskadan Markaziy Amerikagacha bo'lgan hududni egallab oldi.[38][101]

Polineziya

The common starling appears to have arrived in Fiji in 1925 on Ono-i-lau and Vatoa orollar. It may have colonised from New Zealand via Raul ichida Kermadek orollari where it is abundant, that group being roughly equidistant between New Zealand and Fiji. Its spread in Fiji has been limited, and there are doubts about the population's viability. Tonga was colonised at about the same date and the birds there have been slowly spreading north through the group.[106][107]

Janubiy Afrika

In South Africa, the common starling was introduced in 1897 by Sesil Rods. It spread slowly, and by 1954, had reached Klanvilliam va Port Elizabeth. It is now common in the southern Cape region, thinning out northwards to the Johannesburg area. U mavjud G'arbiy Keyp, Sharqiy Keyp va Free State provinces of South Africa and lowland Lesoto, with occasional sightings in KwaZulu-Natal, Gauteng va shaharcha atrofida Oranjemund yilda Namibiya. In Southern Africa populations appear to be resident and the bird is strongly associated with man and anthropogenic habitats. It favours irrigated land and is absent from regions where the ground is baked so dry that it cannot probe for insects. It may compete with native birds for crevice nesting sites but the indigenous species are probably more disadvantaged by destruction of their natural habitat than they are by inter-specific competition. It breeds from September to December and outside the breeding season may congregate in large flocks, often roosting in qamishzorlar. It is the most common bird species in urban and agricultural areas.[108]

G'arbiy Hindiston

In 1901, the inhabitants of Sent-Kits petitioned the Colonial Secretary for a ″government grant of starlings to exterminate″ an outbreak of grasshoppers which was causing enormous damage to their crops.[109] The common starling was introduced to Jamaica in 1903, and the Bahamas and Cuba were colonised naturally from the US.[24][110] This bird is fairly common but local in Jamaica, Katta Bahama va Bimini, and is rare in the rest of the Bahamas, eastern Cuba,[111] The Kayman orollari, Puerto-Riko va Sankt-Croix.[112]

Holat

The global population of the common starling is estimated to be more than 310 million individuals and its numbers are not thought to be declining significantly, so the bird is classified by the Tabiatni muhofaza qilish xalqaro ittifoqi bo'lgani kabi eng kam tashvish.[1] It had shown a marked increase in numbers throughout Europe from the 19th century to around the 1950s and 60s. Taxminan 1830 yilda, S. v. vulgaris expanded its range in the British Isles, spreading into Ireland and areas of Scotland where it had formerly been absent, although S. v. zetlandicus was already present in Shetland and the Outer Hebrides. The common starling has bred in northern Sweden from 1850 and in Iceland from 1935. The breeding range spread through southern France to northeastern Spain, and there were other range expansions particularly in Italy, Austria and Finland.[13] It started breeding in Iberia in 1960, while the spotless starling's range had been expanding northward since the 1950s. The low rate of advance, about 4.7 km (2.9 mi) per year for both species, is due to the suboptimal mountain and woodland terrain. Expansion has since slowed even further due to direct competition between the two similar species where they overlap in southwestern France and northwestern Spain.[15][113]

Major declines in populations have been observed from 1980 onward in Sweden, Finland, northern Russia (Kareliya ) va Boltiqbo'yi davlatlari, and smaller declines in much of the rest of northern and central Europe.[13] The bird has been adversely affected in these areas by intensive agriculture, and in several countries it has been qizil ro'yxatga kiritilgan due to population declines of more than 50%. Numbers dwindled in the United Kingdom by more than 80% between 1966 and 2004; although populations in some areas such as Shimoliy Irlandiya were stable or even increased, those in other areas, mainly England, declined even more sharply. The overall decline seems to be due to the low survival rate of young birds, which may be caused by changes in agricultural practices.[114] The intensiv dehqonchilik methods used in northern Europe mean there is less pasture and meadow habitat available, and the supply of grassland invertebrates needed for the nestlings to thrive is correspondingly reduced.[115]

Odamlar bilan munosabatlar

Benefits and problems

Simlardagi starlings
Congregating on wires in France
Meva yeyayotgan yulduzcha
Feeding on a windfall apple

Since common starlings eat insect pests such as sim qurtlari, they are considered beneficial in northern Eurasia, and this was one of the reasons given for introducing the birds elsewhere. Around 25 million uy qutilari were erected for this species in the former Sovet Ittifoqi, and common starlings were found to be effective in controlling the grass grub Costelytra zealandica Yangi Zelandiyada.[15] The original Australian introduction was facilitated by the provision of nest boxes to help this mainly hasharotlarga qarshi bird to breed successfully,[39] and even in the US, where this is a pest species, the Qishloq xo'jaligi bo'limi acknowledges that vast numbers of insects are consumed by common starlings.[116]

Common starlings introduced to areas such as Australia or North America, where other members of the genus are absent, may affect native species through competition for nest holes. Shimoliy Amerikada, chickadees, nuxatches, qarag'aylar, binafsha martins va boshqalar qaldirg'ochlar ta'sir qilishi mumkin.[101][117] In Australia, competitors for nesting sites include the qip-qizil va eastern rosellas.[118] For its role in the decline of local native species and the damages to agriculture, the common starling has been included in the IUCN Dunyodagi eng yomon invaziv 100 tur ro'yxati.[119]

European, or common, starlings are habitat generalists meaning they are able to exploit a multitude of habitats, nest sites and food sources. This, coupled with them being lowland birds that easily coexist with humans, enables them to take advantage of other native birds, most particularly woodpecker.[120] European starlings are considered aggressive hamma narsa that utilize an open bill probing technique (described in the “Description” section) that gives them an evolutionary advantage over birds that are mevali mevalar.[121] Their aggressive and gregarious behavior in terms of food thus allows them to outcompete native species. Common starlings are also aggressive in the creation of their nest cavities. Often, starlings will usurp a nest site, for example a tree hollow, and fill it rapidly with bedding and contaminants compared to other species, like the native parrots, that use little to no bedding.[122] As cavity nesters, they are able to outcompete many native species in terms of habitat and nest sites.

Common starlings can eat and damage fruit in orchards such as uzum, shaftoli, zaytun, smorodina va pomidor or dig up newly sown grain and sprouting crops.[41][123] They may also eat animal feed and distribute seeds through their droppings. In eastern Australia, weeds like bridal creeper, karapuz va boneseed are thought to have been spread by common starlings.[124] Agricultural damage in the US is estimated as costing about US$800 million annually.[116] This bird is not considered to be as damaging to agriculture in South Africa as it is in the United States.[67]

Common starlings take advantage of agricultural fields, livestock facilities, and other human related sources of food and nest sites. Starlings often assault crops such as grapes, olives, and cherries by consuming excessive amounts of crops in large flock sizes and in new grain fields, starlings pull up young plants and eat the seeds.[125] In caged trials, it was shown that starlings eat 7-23 g of animal food daily and 20-40 g of plant food meaning a decent portion of crops are consumed by these birds.[126] Bird damage to grapes in 1968 cost upwards to $4.4 million while losing almost 17% of the crops.[126] As well, these birds often congregate at feeding troughs to eat grain and concurrently contaminate the food and water source with their droppings that were provided for livestock.[127] For example, high protein supplements added to cattle feed are selectively eaten by common starlings.[128] In 1968, the cost of cattle rations consumed during winter by starlings was $84 per 1,000 starlings and is proposed to be much more expensive today given an increase in current cattle feed costs.[126] The English or house sparrow (Passer domesticus ) and the common starling are considerable agricultural pests, together causing an estimated US$ 1 billion per year in crop damages.[129]

The large size of flocks can also cause problems. Common starlings may be sucked into aircraft jet engines, one of the worst instances of this being an incident in Boston in 1960, when sixty-two people died after a turboprop samolyot flew into a flock and plummeted into the sea at Winthrop Makoni.[130] The large roosts of the common starling pose many safety hazards for aircraft, mainly including the clogging of engines that concurrently shutdown the plane into descent.[131] From the years 1990-2001, 852 incidents of aircraft hazard due to starlings and blackbirds were reported with 39 strikes causing major damage that cost a total of $1,607,317.[126]

Starlings' droppings can contain the fungus Histoplazma kapsulatum, sababi gistoplazmoz odamlarda. At roosting sites this fungus can thrive in accumulated droppings.[15] Boshqa qatorlar mavjud yuqumli kasalliklar that can potentially be transmitted by common starlings to humans,[116] although the potential for the birds to spread infections may have been exaggerated.[101] The spread of disease to livestock is also a concern, possibly more important than starling's effects on food consumption or transmission of disease to humans. The spreading of Gistoplazmoz reported in a Nebraska manufacturing facility saw a loss of 10,000 pigs from the spread of the disease which is valued at $1 million US dollars loss in today's market.[126]

Boshqaruv

Because of the damage they do, there have been attempts to control the numbers of both native and introduced populations of common starlings. Within the natural breeding range, this may be affected by legislation. For example, in Spain, this is a species hunted commercially as a food item, and has a closed season, whereas in France, it is classed as a pest, and the season in which it may be killed covers the greater part of the year. In Great Britain, Starlings are protected under the Wildlife and Countryside Act 1981, which makes it "illegal to intentionally kill, injure or take a starling, or to take, damage or destroy an active nest or its contents". The Wildlife Order in Northern Ireland allows, with a general licence, "an authorised person to control starlings to prevent serious damage to agriculture or preserve public health and safety".[132] This species is migratory, so birds involved in control measures may have come from a wide area and breeding populations may not be greatly affected. In Europe, the varying legislation and mobile populations mean that control attempts may have limited long-term results.[123] Non-lethal techniques such as scaring with visual or auditory devices have only a temporary effect in any case.[24]

Huge urban roosts in cities can create problems due to the noise and mess made and the smell of the droppings. In 1949, so many birds landed on the clock hands of London's Big Ben that it stopped, leading to unsuccessful attempts to disrupt the roosts with netting, repellent chemical on the ledges and broadcasts of common starling alarm calls. An entire episode of Goon shousi in 1954 was a parodiya of the futile efforts to disrupt the large common starling roosts in central London.[133]

Qushlarni boqish uchun starling
Visiting a bird feeder. The adult has a black beak in the winter.

Where it is introduced, the common starling is unprotected by legislation, and extensive control plans may be initiated. Common starlings can be prevented from using nest boxes by ensuring that the access holes are smaller than the 1.5 in (38 mm) diameter they need, and the removal of perches discourages them from visiting qushlarni oziqlantiruvchi vositalar.[101]

Western Australia banned the import of common starlings in 1895. New flocks arriving from the east are routinely shot, while the less cautious juveniles are trapped and netted.[97] New methods are being developed, such as tagging one bird and tracking it back to establish where other members of the flock roost.[134] Another technique is to analyse the DNA of Australian common starling populations to track where the migration from eastern to western Australia is occurring so that better preventive strategies can be used.[135] By 2009, only 300 common starlings were left in Western Australia, and the state committed a further A$400,000 in that year to continue the eradication programme.[136]

In the United States, common starlings are exempt from the Ko'chib yuruvchi qushlar to'g'risidagi qonun, which prohibits the taking or killing of migratory birds.[137] No permit is required to remove nests and eggs or kill juveniles or adults.[101] Research was undertaken in 1966 to identify a suitable avitsitsid that would both kill common starlings and would readily be eaten by them. It also needed to be of low toxicity to mammals and not likely to cause the death of pets that ate dead birds. The chemical that best fitted these criteria was DRC-1339, now marketed as Starlicide.[138] In 2008, the United States government poisoned, shot or trapped 1.7 million birds, the largest number of any nuisance species to be culled.[139] In 2005, the population in the United States was estimated at 140 million birds,[140] around 45% of the global total of 310 million.[1]

The likelihood of starlings to damage the feeding operations is dependent on the number of livestock, favoring areas with more livestock.[141] They also show preference for feed types which were not whole corn but smaller feeds, creating more damage in areas where the feed was smaller.[141] They also showed feed preference based on composition.[142] A proposed solution to this problem is use of less palatable feed by agriculturalists, perhaps relying on larger feed types or feed which is less favorable in composition to starlings.[142][141] An additional solution for mitigation control involves ensuring that livestock feeding operations are not within close proximity of each other or starling roosts.[141] Weather conditions also had an impact on whether starlings visited livestock feeding operations, with a higher likelihood to visit in colder temperatures or following snow storms.[143]

Alternatives to managing starling populations in agricultural areas include the use of starlicide. Use of starlicide has been found to reduce the spread of Salmonella enterica in livestock and other diseases found among livestock.[143] Though this does not appear to eliminate introduction of these diseases completely, it has been determined that they are contributors and starling control is a successful mitigation strategy.[143]

Ilm-fan va madaniyatda

Uy hayvonlari qarayapti
Pet in a cage

Common starlings may be kept as pets or as laboratory animals. Avstriyalik etolog Konrad Lorenz wrote of them in his book Shoh Sulaymonning uzugi as "the poor man's dog" and "something to love",[144] because nestlings are easily obtained from the wild and after careful hand rearing they are straightforward to look after.[144][145] They adapt well to captivity, and thrive on a diet of standard bird feed and ovqat qurtlari. Several birds may be kept in the same cage, and their inquisitiveness makes them easy to train or study. The only disadvantages are their messy and indiscriminate defecation habits and the need to take precautions against diseases that may be transmitted to humans. As a laboratory bird, the common starling is second in numbers only to the uy kabutari.[42]

The common starling's gift for mimicry has long been recognised. In the medieval Welsh Mabinogion, Branwen tamed a common starling, "taught it words", and sent it across the Irish Sea with a message to her brothers, Kepak va Manawydan, who then sailed from Wales to Ireland to rescue her.[146] Katta Pliniy claimed that these birds could be taught to speak whole sentences in Latin and Greek, and in Genri IV, William Shakespeare had Hotspur declare "The king forbade my tongue to speak of Mortimer. But I will find him when he is asleep, and in his ear I'll holler 'Mortimer!' Nay I'll have a starling shall be taught to speak nothing but Mortimer, and give it to him to keep his anger still in motion."

Motsartning
Mozart's "starling song"

Motsart bor edi pet common starling which could sing part of his Piano Concerto in G Major (KV. 453).[133] He had bought it from a shop after hearing it sing a phrase from a work he wrote six weeks previously, which had not yet been performed in public. He became very attached to the bird and arranged an elaborate funeral for it when it died three years later. It has been suggested that his Musiqiy hazil (K. 522) might be written in the comical, inconsequential style of a starling's vocalisation.[37] Other people who have owned common starlings report how adept they are at picking up phrases and expressions. The words have no meaning for the starling, so they often mix them up or use them on what to humans are inappropriate occasions in their songs.[147] Their ability at taqlid is so great that strangers have looked in vain for the human they think they have just heard speak.[37]

Common starlings are trapped for food in some Mediterranean countries.[15] The meat is tough and of low quality, so it is casseroled or made into pâté. One recipe said it should be stewed "until tender, however long that may be". Even when correctly prepared, it may still be seen as an acquired taste.[133][148][149]

Izohlar

  1. ^ The table is based on Feare & Craig (1998).[15] Parentheses indicate that the scientific name has changed from that originally given.
  2. ^ This form was described by Hodgson as S. indicus yilda Kulrang "s Zoologik xilma-xillik of 1831, and may have taksonomik ustuvorlik ustida humii.[16][17]

Adabiyotlar

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